Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanorana quadranus (Liu, Hu, and Yang, 1960)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Nanorana > Species: Nanorana quadranus

Rana quadranus Liu, Hu, and Yang, 1960, Acta Zool. Sinica, 12: 286, 292. Holotype: CIB 571357, by original designation (misprinted in the English abstract as 571589). Type locality: "Kwan-yang, Wushan, Szechwan [=Sichuan], 1463 m altitude", China.

Rana quadrana — Hu, 1985, in Frost (ed.), Amph. Species World: 511. Incorrect subsequent spelling.

Rana (Paa) quadranus — Dubois, 1987 "1986", Alytes, 5: 47.

Paa (Quadrana) quadrana — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 154; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 272.

Chaparana (Feirana) quadranus — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 318.

Rana quadrana — Zhao and Adler, 1993, Herpetol. China: 147.

Paa (Feirana) quadrana — Fei, 1999, Atlas Amph. China: 220.

Nanorana quadranus — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 367.

Gynandropaa (Feirana) quadranus — Ohler and Dubois, 2006, Zoosystema, 28: 781.

Feirana quadranus — Wang, Jiang, Chen, Xie, and Zheng, 2007, Acta Zootaxon. Sinica, 3: 139–145; Wang, Jiang, Xie, Chen, Dubois, Liang, and Wagner, 2009, Zool. Sci., Tokyo, 26: 500–09; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 26. See comment under Dicroglossidae.

Nanorana (Chaparana) quadranus — Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, Suppl. Inform., doi:10.1073/pnas.1008415107/-/DCSupplemental: 2; Hofmann, Baniya, Litvinchuk, Miehe, Li, and Schmidt, 2019, Ecol. Evol., 9: 14506.

English Names

Kwang-yang Asian Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 98).

Swelled Vent Frog (Fei and Ye, 2001, Color Handbook Amph. Sichuan: 188).

Swelled-vented Frog (Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 476).

Distribution

High-gradient streams in forest and shrublands of the Wuling, Daba, Qinling, and Longment mountains of southern Shaanxi, southern Shanxi, southwestern Henan, southern Gansu, northeastern Sichuan, Hubei, northern Hunan, and western Anhui, China, 500–2700 m elevation.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Related to Rana spinosa according to the original publication, although Dubois' taxonomy suggests that this is incorrect. See Liu and Hu, 1961, Tailless Amph. China: 150–153. See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 289; and Fei, 1999, Atlas Amph. China: 220–221. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 188, provided a brief account and illustration (as Paa quadrana). Fan, Guo, and Liu, 1998, Amph. Rept. Shanxi Prov.: 73–75, provided an account for Shanxi. See statement of geographic range, habitat, and conservation status (as Chaparana quadranus) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 628. See comment under Nanorana with regard to the status of Feirana. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1430–1437, provided an accounts (as Feirana quadranus), figures, and map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 382–383, provided a brief account (as Feirana quadranus) including photographs of specimens and habitat. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 476–477, provided an account (as Feirana quadranus), photographs, and a range map. Huang, Duan, and Ji, 2014, Zootaxa, 3779: 497–500, reported on larval morphology (as Feirana quadranus). See comments by Zhao, Wang, Wang, Liu, and Lu, 2015, J. Henan Normal Univ. (Nat. Sci.), 46: 705–711, reassigning previous records from Henan as Feirana taihangnicus and Yerana yei. Yao and Gong, 2012, Amph. Rept. Gansu: 57–58, provided a brief account and photographs. Shen, 2014, Fauna Hunan, Amph.: 294–298, provided an account for Hunan, China. Yang, Jiang, Luo, Yang, Wang, Liao, and Hu, 2019, Asian Herpetol. Res., 10: 48–55, reported on microhabitat segregation between this species and Nanorana taihangnica (as Feirana) in the Qinling Mountains.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.