Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Boulenophrys wushanensis (Ye and Fei, 1995)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Megophryinae > Genus: Boulenophrys > Species: Boulenophrys wushanensis

Megophrys wushanensis Ye and Fei, 1995, Acta Herpetol. Sinica, 4–5: 74–75, 80. Holotype: CIB 571991, by original designation; now renumbered CIB 98012, according to Lyu, Qi, Wang, Zhang, Zhao, Zeng, Wan, Yang, Mo, and Wang, 2023, Zool. Res., Kunming, 44: 441. Type locality: "Miaotang (31° 27′ N, 110° 56′ E), Wushan, Sichuan, altitude 2000 m", China.

Megophrys (Xenophrys) wushanensis — Dubois and Ohler, 1998, Dumerilia, 4: 14.

Xenophrys wushanensis — Ohler, 2003, Alytes, 21: 23, by implication; Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 17; Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 106: 41.

Boulenophrys wushanensis — Fei and Ye, 2016, Amph. China, 1: 670; Qi, Lyu, Wang, Mo, Zeng, Zeng, Dai, Li, Grismer, and Wang, 2021, Zootaxa, 5072: 404.

Xenophrys (Panophrys) wushanensis — Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 109: 41.

Megophrys (Panophrys) wushanensis — Mahony, Foley, Biju, and Teeling, 2017, Mol. Biol. Evol., 34: 755.

Panophrys wushanensis — Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 13.

English Names

Wushan Horned Toad (Fei, 1999, Atlas Amph. China: 122).

Distribution

Known from multiple localities in the Wushan and Daba mountains along the border of Hubei, Chongqing, Sichuan, Shaanxi, and Gansu at elevations of 900–1500 m.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Previously considered to be part of Megophrys minor (e. g, Liu, Hu, and Yang, 1960, Acta Zool. Sinica, 12: 280). Fei, 1999, Atlas Amph. China: 122–123, provided a brief account, figure, and map. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 151, provided a brief account and illustration. In the Megophrys boettgeri group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 82. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 451–455, provided an account, illustration of the holotype, and spot map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 215, provided a brief account including photographs. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 244, provided an account, photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 670–672, provided an account, photographs, and dot map. as Boulenophrys wushanensis. See Yao and Gong, 2012, Amph. Rept. Gansu: 29–30, who provided a brief account and photograph. Lyu, Qi, Wang, Zhang, Zhao, Zeng, Wan, Yang, Mo, and Wang, 2023, Zool. Res., Kunming, 44: 440–441, provided an account and corrected some descriptive errors in the literature.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.