Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Phrynobatrachus cornutus (Boulenger, 1906)

Class: Amphibia > Order: Anura > Family: Phrynobatrachidae > Genus: Phrynobatrachus > Species: Phrynobatrachus cornutus

Arthroleptis cornutus Boulenger, 1906, Ann. Mag. Nat. Hist., Ser. 7, 17: 318. Syntypes: BMNH 1947.2.30.49–50 (formerly 1906.5.28.103–104), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 444, and museum records. Type locality: "Zima" (= Sangmelima, Départment Dja-et-Lobo), Cameroon. See comment by Frétey, Dubois, and Ohler, 2014, Alytes, 30: 44.

Micrarthroleptis cornutus — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 159.

Phrynobatrachus (Micrarthroleptis) cornutus — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 208.

Phrynobatrachus cornutus — Guibé and Lamotte, 1958, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 30: 257; Guibé and Lamotte, 1963, Mem. Inst. Franç. Afr. Noire, 66: 626; Mertens, 1965, Bonn. Zool. Beitr., 16: 22.

Phrynobatrachus cornutus cornutus — Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 365.

English Names

Horned River Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 103).

Horned Puddle Frog (Jacobsen, 2009, Afr. Herp News, 47: 5).

Distribution

Western Cameroon to northwestern Dem. Rep. Congo; Bioko and Equatorial Guinea through Gabon to north-east-central Central African Republic; likely into northwestern Angola.

Geographic Occurrence

Natural Resident: Cameroon, Central African Republic, Congo, Democratic Republic of the, Congo, Republic of the, Equatorial Guinea, Gabon

Likely/Controversially Present: Angola

Comment

Discussed by Perret, 1988, Arch. Sci., Genève, 41: 279–281. See Largen and Dowsett-Lemaire, 1991, Tauraco Res. Rept., 4: 151–152, for Rep. Congo records. Frétey and Blanc, 2002 "2001", Bull. Soc. Zool. France, 126: 386, reported this species from Gabon. Frétey, 2008, Alytes, 25: 99–172, summarized the literature. Jacobsen, 2009, Afr. Herp News, 47: 2–20, reported this species from east-north-central Central African Republic. In species group B of Zimkus, Rödel, and Hillers, 2010, Mol. Phylogenet. Evol., 55: 883–900. Nagy, Chifundera, Collet, and Gvoždík, 2013, Herpetol. Notes, 6: 413–419, provided a record from Bas-Congo, southwestern Dem. Rep. Congo. Baptista, Conradie, Vaz Pinto, and Branch, 2019, In Huntley, Russo, Lages, and Ferrand (eds.), Biodiversity in Angola: 256, noted specimens in Luki, Dem. Rep. Congo, near Angola (both the Cabinda enclave and northwestern Angola proper), suggesting its possible occurrence there. Dewynter and Frétey, 2019, Cah. Fondation Biotope, 27: 33, summarized the literature for Gabon and provided a photograph (p. 61). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 298–299, provided a brief account, photograph, and range map. In the Phrynobatrachus calcaratus group of Goutte, Reyes-Velasco, and Boissinot, 2019, ZooKeys, 824: 53–70. See brief account and range map for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.