Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Kurixalus verrucosus (Boulenger, 1893)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Kurixalus > Species: Kurixalus verrucosus

Rhacophorus verrucosus Boulenger, 1893, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 13: 337. Syntypes: (3 specimens) BMNH and MSNG; BMNH specimen (1893.9.23.16 [reregistered 1947.2.8.91], by museum records) invalidly considered holotype by Boulenger, 1895 "1894", Proc. Zool. Soc. London, 1894: 640–646; MSNG 29853 designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 216. Type locality: "Thao", Myanmar.

Rhacophorus (Rhacophorus) verrucosus — Ahl, 1931, Das Tierreich, 55: 57, 122.

Rhacophorus (Rhacophorus) appendiculatus verrucosus — Wolf, 1936, Bull. Raffles Mus., 12: 162.

Aquixalus (Aquixalus) verrucosus — Delorme, Dubois, Grosjean, and Ohler, 2005, Bull. Mens. Soc. Linn. Lyon, 74: 166.

Kurixalus verrucosus — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 245.

Kurixalus (Aquixalus) verrucosus — Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 56.

English Names

Boulenger's Bushfrog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 154).

Small Rough-armed Tree Frog (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 41).

Boulenger's Tree Frog (Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 406).

Warty Frilled Treefrog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 56).

Small Frilled-limbed Treefrog (Zug, 2022, Smithson. Contrib. Zool., 653: 41).

Distribution

Highlands of central and northern Vietnam, Laos, and Yunnan, Guizhou, Fujian, Hainan I., and Guangxi, China, through western and northern peninsular Thailand (Tak Province to Prachuap Khiri Khan Province) to northern Myanmar (Kachin and Kayah).

Geographic Occurrence

Natural Resident: China, People's Republic of, Laos, Myanmar, Thailand, Vietnam

Comment

Removed from the synonymy of Rhacophorus appendiculatus by Inger, Orlov, and Darevsky, 1999, Fieldiana, Zool., N.S., 92: 39, where it had been placed by Wolf, 1936, Bull. Raffles Mus., 12: 162 (although this synonymy was accepted only provisionally by Inger, 1954, Fieldiana, Zool., 33: 374, and implicitly not accepted by Inger and Tan, 1996, Raffles Bull. Zool., 44: 567). Inger, Orlov, and Darevsky, 1999, Fieldiana, Zool., N.S., 92: 39–40, discussed confusion in the application of names (Rhacophorus cavirostris, Rhacophorus appendiculatus, Rhacophorus verrucosus, and Rhacophorus bisacculus) and suggested that record of Rhacophorus cavirostris from China and Vietnam actually represented this species or Rhacophorus bisacculus. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 678, suggested that at least some records of Rhacophorus or Philautus cavirostris from China applied to Kurixalus odontotarsus. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 98, also discussed the problem of identification in Vietnam. Considerable Chinese literature deals with this frog (or Kurixalus odontotarsus) under the name Rhacophorus cavirostris (e.g., Liu and Hu, 1961, Tailless Amph. China: 247–248, who regarded this as a species distinct from Rhacophorus buergeri microtympanum (= Philautus microtympanum), in which it was placed as a synonym by Wolf, 1936, Bull. Raffles Mus., 12: 137–217. See account of Vietnam population by Orlov, Lathrop, Murphy, and Ho, 2001, Russ. J. Herpetol., 8: 39–40. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 154–155, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 41, provided specific localities for Vietnam. Orlov, Nguyen, and Ho, 2008, Russ. J. Herpetol., 15: 67–84, provided a photograph (as Rhacophorus verrucosus) and a key to differentiate this species from others in Vietnam. Wiens, Sukumaran, Pyron, and Brown, 2009, Evolution, 63: 1217–1231, corroborated the placement of this species in Kurixalus. Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 138, detailed the range in Thailand. Yu, Zhang, and Yang, 2010, Mol. Phylogenet. Evol., 56: 942–950, and Yu, Zhang, and Yang, 2013, Biochem. Syst. Ecol., 47: 31–37, verified the distinctiveness of this species from Kurixalus bisacculus and Kurixalus odontotarsus. Yu, Rao, Matsui, and Yang, 2017, Sci. Rep. (Nature, London), 7 (16124): 1–13, noted that records of Kurixalus verrucosus from Xizang, China, are referable to Kurixalus naso. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 406–407, provided a brief account (description, photographs, habitat, and range) for Thailand (in Thai). Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 160, provided photographs, a brief account for Thailand, and a range map. Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 56, implied that this may be a species complex. Zug, 2022, Smithson. Contrib. Zool., 653: 41, briefly discussed identification, habitat, and range in Myanmar.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.