Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Plethodon wehrlei Fowler and Dunn, 1917

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Plethodontinae > Genus: Plethodon > Species: Plethodon wehrlei

Plethodon wehrlei Fowler and Dunn, 1917, Proc. Acad. Nat. Sci. Philadelphia, 69: 23. Holotype: ANSP 19123, by original designation. Type locality: "Two Lick Hills, Indiana County, Pennsylvania", USA.

Plethodon wehrlei wehrlei — Bogert, 1954, Amph. Rept. World, 2: 1195; Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 234.

Plethodon (Plethodon) wehrlei — Vieites, Nieto-Roman, Wake, and Wake, 2011, Mol. Phylogenet. Evol., 59: 632, by implication.

English Names

Wehrle's Salamander (Plethodon wehrlei: Bishop, 1943, Handb. Salamanders: 281; Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 35; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 175; Conant, 1975, Field Guide Rept. Amph. E. Cent. N. Am., Ed. 2: 277; Collins, Huheey, Knight, and Smith, 1978, Herpetol. Circ., 7: 8; Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 33; Collins, 1997, Herpetol. Circ., 25: 9; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 29; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 14; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 30; Highton, Bonett, and Jockusch, 2017, in Crother (ed.), Herpetol. Circ., 43: 32).

Wehrle's Salamander (Plethodon wehrlei wehrlei [now coextensive with Plethodon wehrlei]: Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 175). Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 233).

Roanoke Salamander (Plethodon wehrlei: Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 175). Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 233;

Distribution

From southwestern New York to southeastern West Virginia, 200 to 1446 m elevation; presumably into adjacent Virginia, far west Maryland, and southeastern Ohio, USA. See comment.

Geographic Occurrence

Natural Resident: United States of America, United States of America - New York, United States of America - Ohio, United States of America - Pennsylvania, United States of America - West Virginia

Likely/Controversially Present: United States of America - Maryland, United States of America - Virginia

Endemic: United States of America

Comment

In the Plethodon wehrlei group of Highton and Larson, 1979, Syst. Zool., 28: 579–599. See accounts by Highton, 1987, Cat. Am. Amph. Rept., 402: 1–3, and Petranka, 1998, Salamand. U.S. Canada: 409–411. See Cupp and Towles, 1983, Trans. Kentucky Acad. Sci., 44: 157–158, for Kentucky record. Pauley and Watson, 2005, in Lannoo (ed.), Amph. Declines: 853–854, provided a detailed account that summarized the biology and conservation literature. Kozak, Weisrock, and Larson, 2006, Proc. R. Soc. London, Ser. B, Biol. Sci., 273: 539–546, provided molecular evidence of shared mtDNA characteristics of one of the populations of Plethodon punctatus. Highton, Hastings, Palmer, Watts, Hass, Culver, and Arnold, 2012, Mol. Phylogenet. Evol., 63: 278–290, noted evidence that suggests that this nominal species contains cryptic species for which names are likely available, one of these being what is now Plethodon dixi. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 394, provided a brief account, photograph, and range map. Kuchta, Brown, and Highton, 2018, Zool. Scripta, 47: 285–299, discussed the molecular phylogeography of the nominal species, demonstrating analytic paraphyly with Plethodon punctatus, and also recognizing Plethodon dixi, formerly a cavernicolous race in southwestern Virginia. See also discussion by Felix, Wooten, Pierson, and Camp, 2019, Zootaxa, 4609: 429–448, who removed Plethodon jacksoni from synonymy and named Plethodon pauleyi from the southern part of the original range, restricting the name Plethodon wehrlei to their "northern wehrlei". Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 393, provided a brief account, photograph, and range map. Raffaëlli, 2022, Salamanders & Newts of the World: 904, provided an account summarizing morphology, life history, population status, and distribution (including a polygon map).

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.