Bufotes balearicus (Boettger, 1880)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufotes > Species: Bufotes balearicus

Bufo variabilis var. balearica Boettger, 1880, Zool. Anz., 3: 642. Syntypes: SMF; SMF 3722 (formerly 1297.1a) designated lectotype by Mertens, 1967, Senckenb. Biol., 48(A): 40. Type locality: "in insulis Balearicis Majorca et Minorca", Spain. Given as "Palma, Mallorca, Balearen", Spain, by Mertens, 1967, Senckenb. Biol., 48(A): 40. See nomenclatural discussion by Novarini and Bonato, 2010, Acta Herpetol., Firenze, 5: 37–62.

Bufo viridis balearicusHemmer, Kadel, and Kadel, 1981, Amphibia-Reptilia, 2: 217; Speybroeck, Beukema, Dufresnes, Fritz, Jablonski, Lymberakis, Martínez-Solano, Razzetti, Vamberger, Vences, Vörös, and Crochet, 2020, Amphibia-Reptilia, 41: 148. 

Bufo balearicusStöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 667.

Pseudepidalea balearicaFrost, 2007, Amph. Spec. World, vers. 5.0, by implication of results of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365, and Stöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 663–689.

Bufo (Bufotes) balearicusDubois and Bour, 2010, Zootaxa, 2447: 25. See comment.

Bufotes balearicusDufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 23. 

Bufotes viridis balearicus — Speybroeck, Beukema, Dufresnes, Fritz, Jablonski, Lymberakis, Martínez-Solano, Razzetti, Vamberger, Vences, Vörös, and Crochet, 2020, Amphibia-Reptilia, 41: 149. 

English Names

Balearic Green Toad (Dufresnes, 2019, Amph. Eur., N. Afr., & Middle East: 77).

Distribution

Italy (except for the northeast), including Sardinia, Corsica (France), and the Balearic Islands of Spain; possibly Malta. See comment

Comment

Removed from the synonymy of Bufotes viridis by Stöck, Moritz, Hickerson, Frynta, Dujsebayeva, Eremchenko, Macey, Papenfuss, and Wake, 2006, Mol. Phylogenet. Evol., 41: 663–689, on the basis of molecular data, who noted this is a diploid species. See also account (as Bufo viridis) of Corsica population by Delaguerre and Cheylan, 1992, Atlas Repart. Batr. Rept. Corse: 33–34. Barbadillo-Escrivá, 1987, Guia INCAFO Anf. Rept. Peninsula Iberica: 159–162, provided an account (as Bufo viridis) for the Spanish populations. Lanza, 1983, Guide Reconoscimento Spec. Animali, Anf. Rett.: 107–110, reported on the Italian populations as Bufo viridis, which presumably are now, except for the northeastern part of the country, assigned to Bufotes balearicus. See comment by Lever, 2003, Naturalized Rept. Amph. World: 176–177, regarding the distribution in the Balearics. El-Oualidi and Jaziri, 2001, Bull. Inst. Scient., Rabat, 23: 1–8, suggested that the population on Corsica is a distinct species (prior to the recognition of Bufotes balearicus). The population of "Bufo viridis" on Malta may be assignable to Bufotes balearicus but does not seem to have been addressed by the appropriate authors (DRF). See comment under Bufotes siculus, which was previously confused with this taxon. See detailed account (as Bufo viridis) by Balletto, Bologna, and Giacoma, 2007, in Lanza et al. (eds.), Fauna d'Italia, 42 (Amph.): 304–305, for Italian population. Rivera, Escoriza, Maluquer-Margalef, Arribas, and Carranza, 2011, Amf. Rept. Catalunya: 81–83, provided a brief account for the Balearic populations. Mateo, Ayres, and López-Jurado, 2011, Bol. Asoc. Herpetol. Esp., 22: 4, discussed what they considered to be ancient introduced populations in the Balearic Islands. Pinya and Carretero, 2011, Acta Herpetol., Firenze, 6: 59–80, regarded the populations on the Balearic Islands to represent ancient introductions. Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 1–25, provided a phylogenetic placement and summary of the knowledge of this diploid species, its range and biogeography. See Dufresnes, 2019, Amph. Eur., N. Afr., & Middle East: 77, for brief summary of identifying morphology and biology, a range map, as well as a photograph. Speybroeck, Beukema, Dufresnes, Fritz, Jablonski, Lymberakis, Martínez-Solano, Razzetti, Vamberger, Vences, Vörös, and Crochet, 2020, Amphibia-Reptilia, 41: 148, discussed their reasons for considering this a subspecies of Bufotes viridis.  

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