Trachycephalus quadrangulum (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Trachycephalus > Species: Trachycephalus quadrangulum

Hyla quadrangulum Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 367. Holotype: BMNH 1947.2.13.75 (formerly 1860.6.16.120) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 94, this specimen figured on plate 25, fig. 2 of the original publication. Type locality: "W. Ecuador".

Trachycephalus quadrangulum — Ron, Venegas, Ortega-Andrade, Gagliardi-Urrutia, and Salerno, 2016, ZooKeys, 630: 143. 

English Names

Chocoan Milk Frog (Armijos-Ojeda, Székely, Székely, Cogǎlniceanu, Cisneros-Heredia, Ordóñez-Delgado, Escudero, and Espinosa, 2021, ZooKeys, 1063: 32). 

Distribution

Pacific coast and west slope of the western Andes of Ecuador from western Esmeraldas Province south and as far east as Los Rios Province, and south to Loja Province; likely to extend into adjacent extreme northwestern Peru. 

Geographic Occurrence

Natural Resident: Ecuador

Likely/Controversially Present: Peru

Endemic: Ecuador

Comment

Removed from the synonymy of Trachycephalus coriaceus by Ron, Venegas, Ortega-Andrade, Gagliardi-Urrutia, and Salerno, 2016, ZooKeys, 630: 143, where it had been placed by Duellman, 1971, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 4: 10 (previously placed in the synonymy of Hyla mesophaea by Condit, 1964, J. Ohio Herpetol. Soc., 4: 94). Armijos-Ojeda, Székely, Székely, Cogǎlniceanu, Cisneros-Heredia, Ordóñez-Delgado, Escudero, and Espinosa, 2021, ZooKeys, 1063: 32, provided a record from near Boaspamba, Loja Province, Ecuador, and provided a dot map for western Ecuador. 

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