Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Lithobates brownorum (Sanders, 1973)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Lithobates > Species: Lithobates brownorum

Rana berlandieri brownorum Sanders, 1973, J. Herpetol., 7: 87. Holotype: SM 12498, by original designation. Type locality: "41 mi. W of Xicalango, Campeche, Mexico".

Rana brownorum — Hillis, 1981, Copeia, 1981: 312–319.

Rana (Rana) brownorum — Dubois, 1987 "1986", Alytes, 5: 41, by implication.

Rana (Pantherana) brownorum — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 331; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Lithobates brownorum — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1–13; by implication.

Lithobates (Lithobates) brownorum — Dubois, 2006, C. R. Biol., Paris, 329: 829; Dubois, 2006, Mol. Phylogenet. Evol., 42: 325.

Common Names

Browns' Leopard Frog (Liner, 1994, Herpetol. Circ., 23: 27).

Brown's Leopard Frog (Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 16).

Browns' Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 106).

Central American Waterfrog (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 15).

Distribution

Southern Veracruz and northeastern Oaxaca east through the Yucatan Peninsula and the uplands of Chiapas to Nicaragua, sea level to 1650 m elevation.

Geographic Occurrence

Natural Resident: Belize, Guatemala, Honduras, Mexico, Nicaragua

Comment

Subgenus Rana, Rana berlandieri subgroup of the Rana pipiens group as defined by Hillis, Frost, and Wright, 1983, Syst. Zool., 32: 132–143. Although tentatively considered a species distinct from Rana berlandieri by Hillis, 1981, Copeia, 1981: 312–319, subsequently authors (e.g., Lee, 1996, Amph. Rept. Yucatan Peninsula: 123, and Campbell, 1998, Amph. Rept. N. Guatemala Yucatan Belize: 71, Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 404) did not accept the distinction but data suggestive of synonymizing them has not been disclosed. Zaldívar-Riverón, León-Regagnon, and Nieto-Montes de Oca, 2004, Mol. Phylogenet. Evol., 30: 38–49, recognized the species on the basis of molecular evidence. McCranie, 2007, Herpetol. Rev., 38: 38, summarized the departmental distribution in Honduras. Köhler, 2011, Amph. Cent. Am.: 291–295, provided a brief summary of natural history and identification key for the species of Lithobates in Central America and provided a range map and photograph for this species.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.