Ingerophrynus galeatus (Günther, 1864)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Ingerophrynus > Species: Ingerophrynus galeatus

English Names

Gamboja Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 41).

Cambodia Toad (Nguyen, Ho, and Nguyen, 2009, Herpetofauna Vietnam: 71).

Bony-headed Toad (Fei, 1999, Atlas Amph. China: 130).

Helmeted Toad (Hendrix, Böhme, and Ziegler, 2009, Herpetol. Notes, 2: 155).


Indochina (montane northern, central and southern Vietnam, Laos, eastern Cambodia), presumably to be found in adjacent Yunnan and Guangxi, China.


See brief account by Bourret, 1942, Batr. Indochine: 179–180. Fei, 1999, Atlas Amph. China: 130–131, provided a brief account, figure, and map. In the Bufo galeatus group according to Hu, Jiang, and Tian, 1984, Acta Herpetol. Sinica, Chengdu, N.S.,, 3 (1): 77–85. Dubois and Ohler, 1999, J. South Asian Nat. Hist., 4: 169, suggested provisional allocation to the Bufo asper group. Liu, Lathrop, Fu, Yang, and Murphy, 2000, Mol. Phylogenet. Evol., 14: 433., suggested that Ingerophrynus galeatus is the sister taxon of the Duttaphrynus melanostictus--Duttaphrynus himalayanus clade, but Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, on the basis of more data placed it within Ingerophrynus. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 17, provided specific localities for Vietnam. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 148, provided a photograph. Stuart, Sok, and Neang, 2006, Raffles Bull. Zool., 54: 136–137, provided an account and record for eastern Cambodia. Stuart, 2005, Herpetol. Rev., 36: 475, provided records for Laos. Population on Hainan Island has now been named as Ingerophrynus ledongensis. Hendrix, Böhme, and Ziegler, 2009, Herpetol. Notes, 2: 155–160, reported on tadpole morphology from a locality in the central highlands of Vietnam. See account, photograph, and map in Vassilieva, Galoyan, Poyarkov, and Geissler, 2016, Photograph. Field Guide Amph. Rept. Lowland S. Vietnam: 59–60. 

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