Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptodactylus syphax Bokermann, 1969

Class: Amphibia > Order: Anura > Family: Leptodactylidae > Subfamily: Leptodactylinae > Genus: Leptodactylus > Species: Leptodactylus syphax

Leptodactylus syphax Bokermann, 1969, Rev. Brasil. Biol., 29: 13. Holotype: WCAB 16141, by original designation; now MZUSP 73851, according to de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 46. Type locality: "São Vicente (Gustavo Dutra), Cuiabá, 600 m, Mato Grosso, Brasil".

English Names

Basin White-lipped Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 82).

Whistling Foam Frog (Eterovick and Sazima, 2004, Anf. Serra do Cipó: 97).

Distribution

Central to northeastern Brazil (Minas Gerais, Goiás, Mato Gross, Piauí, Paraíba, south to São Paulo); southern Paraguay (Cordillera Province); far eastern Bolivia (Santa Cruz).

Geographic Occurrence

Natural Resident: Bolivia, Brazil, Paraguay

Comment

In the Leptodactylus pentadactylus group of Heyer, 1972, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 231: 1-8, and Heyer, 1979, Smithson. Contrib. Zool., 301: 1-43. See Heyer, 1995, Proc. Biol. Soc. Washington, 108: 713-714, for discussion of distribution and Paraguay record. Cardoso and Heyer, 1995, Alytes, 13: 67–76, reported on vocalizations. Köhler, 1995, Herpetofauna, Weinstadt, 17: 6-8, and De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 43, provided the Bolivian citation. Silva and Facure, 2007, Herpetol. Rev., 38: 215, provided a record for Goiás, Brazil, and discussed the range. Brusquetti and Lavilla, 2006, Cuad. Herpetol., 20: 15, briefly discussed the range in Paraguay. Eterovick and Sazima, 2004, Anf. Serra do Cipó: 97-98, provided a photograph and brief account. Silva, Giaretta, and Facure, 2008, Contemp. Herpetol., 1: 1–6, reported on the advertisement call. Martins and Silva, 2009, Check List, 5: 433-435, provided a map of know localities and reported the first records for São Paulo, Brazil. Reviewed by Heyer, Heyer, and de Sá, 2010, Cat. Am. Amph. Rept., 868: 1-9. Andrade, Guimarães, Leite, and Leite, 2011, Check List, 7: 592-593, provided a range extension to Piauí, Brazil, and discussed the range. Weiler, Núñez, Airaldi, Lavilla, Peris, and Baldo, 2013, Anf. Paraguay: 100, provided a brief account, image, and dot map for Paraguay. In the Leptodactylus fuscus species group of de Sá, Grant, Camargo, Heyer, Ponssa, and Stanley, 2014, S. Am. J. Herpetol., 9(Spec. Issue 1): 1–123, and who provided a summary of relevant literature (adult and larval morphology, identification, advertisement call, and range) on pp. 46–47. Schulze, Jansen, and Köhler, 2015, Zootaxa, 4016: 79–80, described, diagnosed, and pictured the larva. Neves, Yves, Pereira Silva, Alves, Vasques, Coelho, and Silva, 2019, Herpetozoa, Wien, 32: 113–123, provided habitat information and records for western Minas Gerais, Brazil. Vaz-Silva, Maciel, Nomura, Morais, Guerra Batista, Santos, Andrade, Oliveira, Brandão, and Bastos, 2020, Guia Ident. Anf. Goiás e Dist. Fed. Brasil Central: 130–131, provided an account for Goiás and the D.F., Brazil. Eterovick, Souza, and Sazima, 2020, Anf. Serra do Cipó: 1–292, provided an account, life history information, and an identification scheme for the Serra de Cipó, Minas Gerais, Brazil.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.