Tomopterna cryptotis (Boulenger, 1907)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna cryptotis

Rana cryptotis Boulenger, 1907, Ann. Mag. Nat. Hist., Ser. 7, 20: 109. Syntypes: BMNH, by original designation, and MCZ 19268 (on exchange from BMNH, according to Barbour and Loveridge, 1946, Bull. Mus. Comp. Zool., 96: 182). Type locality: "Catequero, Ponang Kuma (Dongwenna), and in the Kafitu Swamps", Angola.

Rana delalandii cryptotisLoveridge, 1936, Field Mus. Nat. Hist. Publ., Zool. Ser., 22: 90.

Pyxicephalus delalandii cryptotisPienaar, 1963, Koedoe, 6: 79.

Pyxicephalus delalandei cryptotisPoynton, 1964, Ann. Natal Mus., 17: 96.

Rana (Pyxicephalus) delalandii cryptotisMertens, 1971, Abh. Senckenb. Naturforsch. Ges., 529: 7.

Tomopterna delalandei cryptotisStevens, 1974, Arnoldia, Zimbabwe, 6: 6.

Tomopterna cryptotisPassmore and Carruthers, 1979, S. Afr. Frogs: 120; Poynton and Broadley, 1985, Ann. Natal Mus., 27: 125.

Rana (Tomopterna) cryptotisDubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Tomopterna (Tomopterna) cryptotisDubois, 1987 "1986", Alytes, 5: 56.

English Names

Striped Pyxie (Pienaar, 1963, Koedoe, 6: 79; Wager, 1965, Frogs S. Afr.: 134; Passmore and Carruthers, 1979, S. Afr. Frogs: 120; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 83).

Striped Burrowing Frog (Pienaar, 1963, Koedoe, 6: 79; Passmore and Carruthers, 1979, S. Afr. Frogs: 120).

Common Burrowing Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15).

Tremolo Sand Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 120; Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 321-322).

Striped Sand Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5).

Catequero Bullfrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Cryptic Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 329).

Tremolo Sand Frog (Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 434).

Common Sand Frog (Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 194).

Distribution

Southern Angola and eastern Namibia east to northeastern Rep. South Africa and Zimbabwe, into southern Zambia and western Mozambique. 

Comment

See Tomopterna milletihorsini for North and East African records previously assigned to this species. See Carruthers and Carruthers, 1979, J. Herpetol. Assoc. Afr., 20: 3-5, for distribution records. See Lanza, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 168, and Poynton and Broadley, 1985, Ann. Natal Mus., 27: 125-127, for account and synonymy. See Salvador, 1996, Smithson. Herpetol. Inform. Serv., 109, for account. Rödel, 2000, Herpetofauna W. Afr., 1: 76-78, provided an account. Pickersgill, 2007, Frog Search: 118-121, provided an account and suggested that this nominal species is composed of several lineages. See Largen, 2001, Tropical Zool., 14: 330-331, for comments on distribution.Channing and Howell, 2006, Amph. E. Afr.: 329-330, provided an account for East Africa, which presumably would not be both Tomopterna cryptotis and Tomopterna milletihorsini. Padial and De la Riva, 2004, Rev. Esp. Herpetol., 18: 95, provided records from Mauritania (now presumably reassigned to Tomopterna milletihorsini, DRF). Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 321-322, provided an account for South Africa. Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 434-435, provided an account for southern Africa. Bates and Haacke, 2003, Navors. Nas. Mus. Bloemfontein, 19: 124-125, discussed the species in Lesotho. See account, photograph, and map for Ethiopia by Largen and Spawls, 2010, Amph. Rept. Ethiopia Eritrea: 194-196 (these records presumably now reassigned to Tomopterna kachowskii, DRF). Zimkus and Larson, 2011, Zootaxa, 2933: 27-45, provided molecular data showing that records from Eritrea, Ethiopia, Djibouti, and Somalia belong to other species (presumably Tomopterna kachowskii). Mercurio, 2011, Amph. Malawi: 291, provided an account for Malawi. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 356–357, provided information on comparative larval morphology. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 153–154, provided a map for Angola and brief discussion of the literature. Phaka, Netherlands, Kruger, and Du Preez, 2017, Bilingual Field Guide Frogs Zululand: 71, provided a photograph, Zululand regional map, and a brief account of life history and identification. Wilson and Channing, 2019, Zootaxa, 4609: 225-246, provided a map that should help with the confusion in the earlier literature. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 378–379, provided a brief account, photograph, and range map. Baptista, António, and Branch, 2019, Amph. Rept. Conserv., 13 (e203): 96–130, reported on specimens from Bicuar National Park, southwestern Angola and commented on the unsettled state of systematics. 

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