Tomopterna milletihorsini (Angel, 1922)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna milletihorsini

Arthroleptis milleti-horsini Angel, 1922, Bull. Mus. Natl. Hist. Nat. Paris, 28: 41. Holotype: MNHNP 1921.635 by original designation; not mentioned by Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat., and regarded as lost by Ohler and Frétey, 2008, Alytes, 25: 173–175. Type locality: "la région caillouteuse de Bélédougou, à Kati (12 kilomètres au nord de Bamakko)", Mali.

Arthroleptis agadezi Angel, 1936, Bull. Soc. Zool. France, 61: 275. Syntypes: MNHNP 36.124-125 (2 specimens), according to the original publication. Type locality: "Agadez (Aïr)", Niger. Considered a junior synonym of Pyxicephalus delalandii (Tschudi) (Somali specimens of this nominal species) by Angel and Guibé, 1948, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 20: 62, before the firm distinction of Tomopterna cryptotis and Tomopterna delalandii. Treated by implication as a synonym of Tomopterna milletihorsini by Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 397.

Schoutedenella milletihorsiniLaurent, 1954, Ann. Mus. R. Congo Belge, Tervuren, Ser. Quarto, Sci. Zool., 1: 40; Laurent, 1985, In Frost (ed.), Amph. Species World: 19.

Arthroleptis milletihorsiniLaurent, 1957, Rev. Zool. Bot. Afr., 56: 274-275, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 336.

Tomopterna milletihorsiniOhler and Frétey, 2008, Alytes, 25: 174.

English Names

Mali Screeching Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 37).

Horsin's Sand Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 382).


North and East subsaharan Africa from Mauretania and Guinea east through northern Nigeria, northern Cameroon, Burkina Faso, northern Central African Rep., southern Sudan and South Sudan to the Ethiopian border, thence south through Kenya and northern Tanzania. 


Ohler and Frétey, 2008, Alytes, 25: 173–175, discussed the identity of the type and the type locality. Zimkus and Larson, 2011, Zootaxa, 2933: 27–45, provided the Nara, Mali record. See detailed account for Egypt by Escoriza and Ben Hassine, 2019, Amph. N. Afr.: 281–236. See comment under Tomopterna cryptotis.  Padial, 2003, Herpetol. Rev., 34: 260, reported the species (as Tomopterna cryptotis) for Mauritania. Spawls, Wasonga, and Drewes, 2019, Amph. Kenya: 35, provided a range map for Kenya, photograph, and brief characterization, as Tomopterna cryptotisSchiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 30, provided records for Nigeria (as Tomopterna delalandii). Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 382–383, provided a brief account, photograph, and range map, as Tomopterna cf. milletihorsini

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