Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eupsophus calcaratus (Günther, 1881)

Class: Amphibia > Order: Anura > Family: Alsodidae > Genus: Eupsophus > Species: Eupsophus calcaratus

Cacotus calcaratus Günther, 1881, Proc. Zool. Soc. London, 1881: 19. Holotype: BMNH 1868.9.22.8 by museum records, according to Lavilla, Nuñez, Rabanal, Langone, and de Sá, 2010, Zootaxa, 2561: 56; now registered 1947.2.20.1, according to museum records. Type locality: "Chiloé" Island, Chile.

Borborocoetes calcaratus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 256.

Zachaenus roseus Cope, 1890 "1889", Proc. U.S. Natl. Mus., 12: 142. Holotype: USNM 15126 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 81. Type locality: "Port Otway, Patagonia"; corrected to "Puerto Almirante Barroso (= Port Otway), 46°49′–75°21′, Península de Taitao, Chile" by Lavilla, Nuñez, Rabanal, Langone, and de Sá, 2010, Zootaxa, 2561: 49–58, who made the synonymy.

Eupsophus calcaratus — Capurro-S., 1958, Invest. Zool. Chilen., 4: 293; Formas and Vera, 1982, Proc. Biol. Soc. Washington, 95: 594.

Common Names

Chiloe Island Ground Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 80; IUCN, 2019).

Rosy Bug-eyed Frog (Zachaenus roseus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 85).

Distribution

Southern Chile and adjacent Argentina (provinces of Chubut, Río Negro, and Neuquen).

Geographic Occurrence

Natural Resident: Argentina, Chile

Comment

Discussed by Formas and Vera, 1982, Proc. Biol. Soc. Washington, 95: 594–601. In the Eupsophus vertebralis group. Synonymy (with Eupsophus grayii) by Cei, 1960, Breviora, 118: 1–13, and Cei, 1962, Invest. Zool. Chilen., 8: 33. Synonymy with Eupsophus roseus by Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 128; Gorham, 1974, Checklist World Amph.: 60, and Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84. Christie and Úbeda, 1996, Herpetol. Rev., 27: 149, provided a new record for Argentina and noted that previous records were based on Eupsophus roseus. Úbeda, 2000, Cuad. Herpetol., 14: 71–74, discussed the distribution in Argentina. Díaz-Páez and Berrios, 2003, Herpetol. Rev., 34: 380, provided a record for Isla Berta and Isla Juan Stuven, Region de Aysen, Chile. Márquez, Penna, Marques, and Amaral, 2005, Herpetol. J., 15: 257–263, reported on advertisement call. Vera Candioti, Úbeda, and Lavilla, 2005, J. Morphol., 262: 161–177, reported on larval morphology. Nuñez, Wood, Rabanal, Fontanella, and Sites, 2011, Mol. Phylogenet. Evol., 58: 343–352, discussed mtDNA phylogeography. See Correa-Quezada and Durán, 2019, ZooKeys, 863: 107–152, for a discussion of systematics and range. See Charrier, 2019, Anf. Bosques Centro Sur y Patagonia de Chile: 170–173, for account (morphology, habitat, natural history) and polygon distribution map.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.