- What is Amphibian Species of the World?
- How to cite
- How to use
- Structure of the taxonomic records
- Running log of additions and corrections, 2024
- Logs of changes and additions, 2014–2023
- What is the right name?
- Curator's blog
- History of the project, 1980 to 2024
- Comments on amphibian taxonomy relating to versions 3.0 to 6.2 (2004 to 2024)
- Scientific Nomenclature and its Discontents: Comments by Frost on Rules and Philosophy of Taxonomy, Ranks, and Their Applications
- Contributors, online editions
- Contributors and reviewers for Amphibian Species of the World: A Taxonomic and Geographic Reference (1985)
- Versions
- Museum abbreviations
- Links to useful amphibian systematic, conservation, collection management, informational, and/or regional sites
- Links to useful FREE library sites
- Copyright and terms of use
Contributors and reviewers for Amphibian Species of the World: A Taxonomic and Geographic Reference (1985)
Angel Alcala
Reviewed: Philippine species.
Address: Department of Biology, Silliman University, Dumaguete City, Philippines.
J.-L. Amiet
Contributed: Cardioglossa (Arthroleptinae) and former Astylosterninae (now part of Arthroleptinae, Arthroleptidae), Petropedetes (now in Petropedetidae) and Phrynodon (now in Phrynobatrachus, Phrynobatrachidae).
Reviewed: Anuran species of Cameroon (except for Arthroleptinae).
Address: Laboratoire de Zoologie, Université de Yaoundé, B.P. 812, Yaoundé, Cameroon.
Steven C. Anderson
Reviewed: Species of southwestern Asia (Turkey to Arabian Peninsula, east through Caucasus region to Pakistan, Afghanistan, and adjacent Russia).
Address: Department of Biological Sciences, College of the Pacific, Stockton, California 95211, USA.
Rose M. A. Blommers-Schlösser
Contributed: Cophylinae, Dyscophus, and Scaphiophryninae (Microhylidae), Mantellinae (now Mantellidae), and Madagascan Rhacophoridae.
Reviewed: Madagascan species of Hyperoliidae, Microhyla (Microhylinae), and Raninae.
Address: Herenstraat 102, 3911 JH Rhenen, Netherlands.
Werner C. A. Bokermann [Deceased 1 April 1995]
Reviewed: Brazilian species.
Address: Parque Zoologico de São Paulo, C.P. 12954, 01000 São Paulo, S.P., Brazil.
Leo J. Borkin
Contributed: Palearctic Raninae (Ranidae) (with some overlap with S. Hu and M. Matsui).
Reviewed: Raninae and species of Palearctic Region, including all of China.
Address: Department of Herpetology, Zoological Institute, Russian Academy of Sciences 1, St. Petersburg 199164, Russia.
Arden H. Brame, Jr. [deceased 19 August 2004]
Contributed: Caudata.
Address: Rosemead, California, USA.
Edmund D. Brodie, Jr.
Reviewed: Caudata.
Address: Department of Biology, UTA Box 19489, University of Texas at Arlington, Arlington, Texas 76019, USA. [Current address: Department of Biology, Utah State University, Logan, Utah 84322-5305, USA.]
Walter C. Brown [deceased]
Contributed: Batrachylodes, Ceratobatrachus, Discodeles, Palmatorappia, and Platymantis (now in Ceratobatrachidae).
Reviewed: Philippine species.
Address: Department of Herpetology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, USA.
Jonathan A. Campbell
Reviewed: Mexican and Central American species.
Address: Department of Biology, UTA Box 19498, University of Texas at Arlington, Arlington, Texas 76019, USA.
David C. Cannatella
Contributed: Centrolenidae (with W.E. Duellman), Leptodactylidae (sensu lato; excluding Brachycephaloidea), and Heleophrynidae.
Reviewed: New World Bufonidae, Phyllomedusinae (Hylidae), and higher classification.
Address: Museum of Natural History, The University of Kansas, Lawrence, Kansas 66045, USA. [Current address: Section of Integrative Biology, The University of Texas, Austin, Texas 78712-1064, USA.]
Ulisses Caramaschi
Reviewed: Brazilian species.
Address: Sector de Herpetologia—Departamento Vertebrados, Museu Nacional do Rio de Janeiro/UFRJ, Quinta da Boa Vista, s/no., 20.942 Rio de Janeiro, R.J., Brazil.
Gustavo Casas-Andreu
Reviewed: Mexican species.
Address: Instituto de Biología, Universidad Nacional Autónoma de México, Apartado Postal 70-153, México 20, D.F., Mexico.
Alan Channing
Contributed: Strongylopus and African Tomopterna (formerly in Ranidae, now Pyxicephalidae).
Reviewed: South African species.
Address: Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X17, Bellville 7530, Rep. South Africa.
Barry T. Clarke
Contributed: Discoglossidae [now Alytidae and Bombinatoridae] and Leiopelmatidae.
Address: formerly Department of Zoology, The Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom.
Ronald I. Crombie
Reviewed: New World Anura.
Address: Division of Amphibians and Reptiles, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560, USA. [Currently Department of Herpetology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, USA.]
J.C. Daniel
Reviewed: Species of India and Sri Lanka.
Address: Bombay Natural History Society, Hornbill House, Shaheed Bhagat Singh Road, Mumbai 400 023, India.
Margaret M. Davies
Reviewed: Species of Australia and New Guinea.
Address: Department of Zoology, University of Adelaide, G.P.O. Box 498, Adelaide, South Australia 5001, Australia. [Currently living in Sandy Bay, Tasmania, Australia.]
Robert C. Drewes
Reviewed: Hyperoliidae.
Address: Department of Herpetology, California Academy of Sciences, Golden Gate Park, San Francisco, California 94118, USA.
William E. Duellman
Contributed: Centrolenidae (with D.C. Cannatella) and Hylidae.
Reviewed: All Amphibia.
Address: Museum of Natural History and Biodiversity Research Center, The University of Kansas, Lawrence, Kansas 66045, USA.
Sushil K. Dutta
Contributed: Ranidae (now Ranidae, Dicroglossidae, and Nyctibatrachidae) of southern Asia (with some overlap with S. Hu and M. Matsui).
Reviewed: Species of India and Sri Lanka.
Address: Museum of Natural History, the University of Kansas, Lawrence, Kansas 66045, USA. [Current address: P.G. Department of Zoology, Utkal University, Vani Vihar, Bhubansewar 751004, Orissa, India]
Stephen R. Edwards
Contributed: Vision, confidence, energy, and money to make the published project a reality.
Address: Association of Systematics Collections, Museum of Natural History, University of Kansas, Lawrence, KS 66045. [Now residing in northeastern Oregon.]
Oscar Flores-Villela
Reviewed: Species of Mexico.
Address: Museo de Zoología, Facultad de Ciencias, Universidad Nacional Autónoma de México, Apartado Postal 70-399, México, D.F. 04510, Mexico.
Linda S. Ford
Reviewed: Dendrobatidae (now Aromobatidae and Dendrobatidae) and Central American brachycephaloids.
Address: Museum of Natural History, The University of Kansas, Lawrence, Kansas 66045, USA. [Current address: Museum of Comparative Zoology, Harvard University, 6 Oxford Street Cambridge, MA 02138.]
Ramón Formas
Contributed: Rhinoderma (now in Rhinodermatidae).
Reviewed: Alsodes and Eupsophus (now Alsodidae), Batrachyla and Hylorhina (now Batrachylidae), Insuetophrynus (now in Rhinodermatidae) and Caudiverbera (now Calyptocephala in Calyptocephalallidae) and Telmatobufo (now in Calyptocephalellidae).
Address: Instituto de Zoología, Universidad Austral de Chile, Casilla de Correo 567, Valdivia, Chile.
Darrel R. Frost
Contributed: All taxa not provided by others, most keystroking, most comments, all library verification of all contributions and all type localities included within quotations, and checking and reconciliation of all contributions and reviews.
Reviewed: All Amphibia.
Address: Museum of Natural History, The University of Kansas, Lawrence, Kansas 66045, USA. [Current address: Division of Vertebrate Zoology (Herpetology), American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024, USA].
Alice G.C. Grandison [deceased 10 March 2014]
Contributed: African Bufonidae and Phrynobatrachus (now in Phrynobatrachidae). With Marinus Hoogmoed the real heavy-lifters who aided DRF in so many ways that I will be eternally grateful.
Address: Department of Zoology, Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom. [Current Address: Grunnadal, Nisbe Rd., Gullane, East Lothian, EH31 2BQK, United Kingdom]
Eduardo Gudynas
Contributed: Pseudopaludicola.
Reviewed: Species of Uruguay, northeastern Argentina, and southern Brazil.
Address: Centro Educativo Don Orione, Departamento de Biología, Casilla de Correo 13125, Montevideo, Uruguay.
Keith A. Harding
Reviewed: Ascaphus (Ascaphidae), New World Bufonidae, Centrolenidae, Dendrobatidae (then including Aromobatidae), New World Microhylidae, New World Pipidae, Pseudis and Lysapsus (Hylidae), Ambystomatidae, Amphiumidae, Plethodontidae, Proteidae, and Sirenidae.
Address: 5 Drakewalls Place, Gunnislake, Cornwall PL18 9EJ, United Kingdom.
W. Ronald Heyer
Reviewed: Leptodactylidae (now composed of Batrachylidae, Calyptocephalellidae, Ceratophryidae, Cycloramphidae, Hylodidae, Leiuperidae, Leptodactylidae, and Telmatobiidae).
Address: Division of Amphibians and Reptiles, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560, USA.
Richard Highton
Reviewed: Salamanders of USA.
Address: Department of Zoology, University of Maryland, College Park, Maryland 20742, USA.
David M. Hillis
Contributed: Nearctic Bufo (now Anaxyrus, Incilius, and Rhinella; Bufonidae) and New World Rana (now Rana and Lithobates; Ranidae; note that DMH disagrees with this taxonomy).
Address: Museum of Natural History, The University of Kansas, Lawrence, Kansas 66045, USA. [Current address: Section of Integrative Biology, The University of Texas, Austin, Texas 78712-1064, USA.]
Marinus S. Hoogmoed
Contributed: South and Central American Bufonidae and Adelophryne and Phyzelaphryne (now in Eleutherodactylidae). With Alice G.C. Grandison the real heavy-lifters who aided DRF in so many ways that he wil be eternally grateful.
Reviewed: All Amphibia.
Address: Nationaal Natuurhistorisch Museum, Postbus 9517, 2300 RA Leiden, Netherlands. [current address: Museu Paraense Emílio Goeldi, Caixa Post 399, 66.000 Belém, Brazil]
Kim M. Howell
Reviewed: Species of East Africa.
Address: Department of Zoology, Box 35064, University of Dar es Salaam, Dar es Salaam, Tanzania.
Shuqin Hu [Deceased 1992]
Contributed: Chinese Raninae (with L. J. Borkin and M. Matsui).
Reviewed: Chinese species.
Address: Chengdu Institute of Biology, Academia Sinica, P.O. Box 416, Chengdu, Sichuan, P. R. China.
Robert F. Inger [deceased 12 April 2019]
Contributed: Asian Bufonidae, Microhylinae, and Rhacophoridae.
Reviewed: Megophryidae.
Address: Division of Amphibians and Reptiles, Field Museum of Natural History, Roosevelt Road at Lake Shore Drive, Chicago, Illinois 60605, USA.
Rafael Joglar
Reviewed: West Indian Eleutherodactylus (now in Eleutherodactylidae and Craugastoridae).
Address: Museum of Natural History, The University of Kansas, Lawrence, Kansas 66045, USA. [Current address: Departamento de Biología, Universidad de Puerto Rico, Río Piedras, Puerto Rico 00931, USA.]
Enrique La Marca
Reviewed: Dendrobatidae (including what is now Aromobatidae) and Venezuelan species of Anura.
Address: School of Life Sciences, University of Nebraska, Lincoln, Nebraska 68508, USA. [Current Address: Calle 15 con Carrera 7ma, Edificio La Marca, Apto. 1, Guanare, Portuguesa 3310, Venezuela.]
Raymond Laurent [deceased 3 February 2005]
Contributed: Arthroleptini (excluding Cardioglossa) (Arthroleptidae), Hemisotidae, and Hyperoliinae (Hyperoliidae).
Reviewed: Species of Africa and higher classification.
Address: Instituto Miguel Lillo, Miguel Lillo 205, Tucumán 4000, Argentina.
Esteban O. Lavilla
Reviewed: Alsodes and Telmatobius (Ceratophryidae).
Address: Instituto Miguel Lillo, Miguel Lillo 205, Tucumán 4000, Argentina.
John D. Lynch
Contributed: Eleutherodactylus (excluding West Indian species and including what is now most of Brachycephalidae, Eleutherodactylidae, and Craugastoridae).
Reviewed: Higher taxonomy.
Address: School of Life Sciences, University of Nebraska, Lincoln, Nebraska 68508, USA. [Current Address: Instituto de Ciencias Naturales, Universidad Nacional de Colombia, A.A. 7495, Bogotá, D.C., Colombia]
Masafumi Matsui
Contributed: Megophryidae and East Asian Raninae (overlapping with L.J. Borkin, S. Dutta, and S. Hu) (Ranidae). Reviewed: East Asian species.
Address: Department of Zoology, Kyoto University, Yoshida, Sakyo, Kyoto 606-01, Japan.
Roy W. McDiarmid
Reviewed: Brachycephalus (Brachycephalidae) and Neotropical Bufonidae.
Address: USGS Patuxent Wildlife Research Center, National Museum of Natural History, Washington, D.C. 20560, USA.
Janre Mullins
Contributed: editorial and clerical and computer support far exceeding paid time.
Address: Association of Systematics Collection, Museum of Natural History, University of Kansas, Lawrence, KS 66045. [Current address:
Financial Analyst, Maine Department of Health and Human Services, 221 State Street, Augusta, ME 04333. email: Janre.Mullins AT maine.gov]
Craig E. Nelson
Contributed: New World Microhylidae.
Address: Department of Biology, Jordan Hall 138, University of Indiana, Bloomington, Indiana 47405, USA.
Ronald A. Nussbaum
Contributed: Sooglossidae.
Reviewed: Gymnophiona.
Address: Museum of Zoology, The University of Michigan, Ann Arbor, Michigan 48104, USA.
Jean-Luc Perret
Contributed: African Raninae (Ranidae) and Dicroglossidae (excluding Strongylopus and Tomopterna).
Reviewed: Hyperoliidae and Petropedetes (Petropedetidae).
Address: Museum d'Histoire Naturelle, Route de Malagnou, C.P. 434, 1211 Geneve 6, Switzerland.
John C. Poynton
Reviewed: African species.
Address: Department of Biological Sciences, University of Natal, King George V Avenue, Durban, Natal 4001, Rep. South Africa [Current address: Department of Zoology, The Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom].
William F. Pyburn [deceased 5 May 2007]
Reviewed: Centrolenidae, New World Hylidae, and New World Microhylidae (Microhylidae).
Address: Department of Biology, UTA Box 19489, University of Texas at Arlington, Arlington, Texas 76019, USA.
Juan A. Rivero [deceased 3 March 2014]
Reviewed: Atelopus (Bufonidae), Dendrobatidae (including Aromobatidae), and South American "Eleutherodactylus" (now Brachycephalidae and Craugastoridae).
Address: Departamento de Biología, Universidad de Puerto Rico, Mayagüez, Puerto Rico 00708, USA.
Jay M. Savage
Reviewed: Centrolenidae, Dendrobatidae (including Aromobatidae), nomenclature, higher taxonomy, and Central America species.
Address: Department of Biology, University of Miami, P.O. Box 249118, Coral Gables, Florida 33124, USA. [Current address: Rana Dorada Enterprises, S.A., 3401 Adams Ave., Suite A, San Diego, CA 92116-2490, USA.]
Arne Schiøtz [deceased 2019]
Reviewed: Hyperoliidae and African Chiromantis (Rhacophoridae).
Address: Danmarks Akvarium, 2920 Charlottenlund, Denmark. [Current address: Humlehaven 2, Atterup, Grevinge, DK-4571 Denmark.]
Albert Schwartz [Deceased 18 October 1992]
Contributed: Peltophryne (Bufonidae) and West Indian "Eleutherodactylus" (now in Craugastoridae and Eleutherodactylidae).
Reviewed: West Indian species.
Address: Miami-Dade Community College, North Campus, Miami, Florida 33167, USA.
Norman J. Scott, Jr.
Reviewed: Central and South American species.
Address: Denver Wildlife Research Center, U.S. Fish and Wildlife Service, Museum of Southwestern Biology, University of New Mexico, Albuquerque, New Mexico 87131, USA. [Current address: 3655 Lindquist Lane, P.O. Box 307, Creston, California 93432. USA.] email address: reptile AT tcsn.net.
Richard Thomas
Reviewed: West Indian species.
Address: Departamento de Biología, Universidad de Puerto Rico, Río Piedras, Puerto Rico 00931, USA.
R.C. Tinsley
Contributed: African Pipidae.
Address: School of Biological Sciences, Queen Mary College (University of London), London E1 4NS, United Kingdom.
Michael J. Tyler [deceased, 26 March 2020]
Contributed: Myobatrachidae (then including Limnodynastidae).
Reviewed: Species of Australia and New Guinea.
Address: Department of Zoology, University of Adelaide, G.P.O. Box 498, Adelaide, South Australia 5001, Australia.
Paulo E. Vanzolini [deceased 28 April 2013]
Reviewed: Brazilian species and nomenclature of higher taxa.
Address: Museu de Zoologia/USP, C.P. 7172, 01.051 São Paulo, S.P., Brazil.
David B. Wake [deceased 29 April 2021]
Reviewed: Caudata.
Address: Museum of Vertebrate Zoology, University of California, Berkeley, California 94720, USA.
Marvalee H. Wake
Contributed: Gymnophiona.
Address: Department of Integrative Biology and Museum of Vertebrate Zoology, University of California, Berkeley, California 94720, USA.
Ermi Zhao [deceased, 24 December 2016]
Reviewed: Species of China.
Address: Chengdu Institute of Biology, Academia Sinica, P.O. Box 416, Chengdu, Sichuan, P. R. China.
Richard G. Zweifel [deceased 25 November 2019]
Contributed: Asterophryinae and Genyophryninae (now all in Asterophryinae: Microhylidae) and original list of current names of living amphibians of the world.
Reviewed: Species of New Guinea.
Address: Department of Herpetology, American Museum of Natural History, Central Park West at 79th Street, New York, New York 10024, USA.