Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eupsophus emiliopugini Formas, 1989

Class: Amphibia > Order: Anura > Family: Alsodidae > Genus: Eupsophus > Species: Eupsophus emiliopugini

Eupsophus emiliopugini Formas, 1989, Proc. Biol. Soc. Washington, 102: 568. Holotype: IZUA 1587, by original designation; destroyed by fire on 3 December 2007. Type locality: "La Picada, 23 km NE (by road) of Ensenada..., Osorno Province, Andean Range, 41°04′ S, 72°26′ W, 480 m.", Chile.

English Names

Emilio's Ground Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 80).

Distribution

Known from the Región Lagos south through to the Región de Aysén (Puerto Yates, Isla Riveros, and including the islands of Chiloé, Kent, and Clemente), southern Chile, and Lago Puelo National Park, Chubut, Argentina.

Comment

In the Eupsophus vertebralis group of Formas and Brieva, 1994, Proc. Biol. Soc. Washington, 107: 391-397. Argentina record from Úbeda, Ramilo, Chehebar, and Vidoz, 1999, Herpetol. Rev., 30: 230. Díaz-Páez and Nuñez, 2002, Herpetol. Rev., 33: 220, provided additional records for Chile. Vera Candioti, Nuñez, and Úbeda, 2010, Acta Zool., Stockholm, 92: 27–45, described development, morphology, and anatomy of the tadpoles. Correa-Quezada and Durán, 2019, ZooKeys, 863: 107–152, discussed the species and noted that its mtDNA phylogeny is not exclusive with respect to Eupsophus vertebralis. Suárez-Villota, Quercia, and Nuñez, 2018, J. Genomics, 6: 98–102, reported on the mitochondrial genome. See Correa-Quezada and Durán, 2019, ZooKeys, 863: 107–152, for discussion of systematics and range. Raimilla and de la Harpe, 2019, Bol. Chil. Herpetol., 6: 59–63, reported localities on Clemente Island, National Reserve Las Guaitecas in the Aysen region, Chile. Quercia, Suárez-Villota, Foresti, and Nuñez, 2020, Comp. Cytogenet., 14: 61–74, discussed the comparative cytogenetics of this species and Eupsophus vertebralis.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.