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* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
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AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
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Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
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Eupsophus roseus (Duméril and Bibron, 1841)

Class: Amphibia > Order: Anura > Family: Alsodidae > Genus: Eupsophus > Species: Eupsophus roseus

Cystignathus roseus Duméril and Bibron, 1841, Erp. Gen., 6: 414. Syntypes: MNHNP 762 (2 specimens), according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 27. Type locality: "Chili" (= Chile). Considered to possibly be Valdivia, Chile, by Cei, 1962, Batr. Chile: 81.

Borborocoetes bibronii Bell, 1843, Zool. Voyage Beagle, Part 5: 35. Holotype: Not designated, although noted as being in the BMNH by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 255; BMNH 1845.5.25.43 (now reregistered 1947.2.19.96), by museum records. Type locality: "Valdivia and Chiloe, in thick forest", Chile. Restricted to "Chonos", Chile by Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 129. Synonymy (with Eupsophus grayii) by Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 128; Cei, 1962, Invest. Zool. Chilen., 8: 33. Synonymy by Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84. Previously placed by Gorham, 1966, Das Tierreich, 85: 118, as a synonym of Eupsophus roseus.

Borborocoetes grayii Bell, 1843, Zool. Voyage Beagle, Part 5: 36. Holotype: Not stated although noted by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 254, as being the in BMNH; BMNH 1947.2.17.83 (formerly 45.5.25.25) according to musem records. Type locality: "in the forest, in Valdivia", Chile. Synonymy by Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 128.

Cystignathus sylvestris Tschudi, 1845, Arch. Naturgesch., 11: 167. Type: Not designated (and not located in MHNN by Peters, 1873, Monatsber. Preuss. Akad. Wiss. Berlin, 1873: 623, so presumed lost). Type locality: "Republica Peruana". Synonymy by Peters, 1873, Monatsber. Preuss. Akad. Wiss. Berlin, 1873: 622; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 253 (tentatively). Regarded as a nomen dubium by Lynch, 1972, Bull. S. California Acad. Sci., 71: 7.

Cystignathus grayi — Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 30.

Cystignathus bibronii — Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 31.

Borborocoetes roseus — Cope, 1865, Nat. Hist. Rev., N.S., 5: 113; Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 36.

Cyclorhamphus fasciatus Peters, 1869, Monatsber. Preuss. Akad. Wiss. Berlin, 1869: 789. Syntypes: ZMB 6617 (2 specimens) according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 45. Type locality: "Puerto Montt (Chile)". Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 257, noted that this species probably belonged in Borborocoetes. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy b Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 128; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84. Previously placed by Gorham, 1966, Das Tierreich, 85: 118, as a synonym of Eupsophus roseus.

Cystignathus fasciatus — Peters, 1870 "1869", Monatsber. Preuss. Akad. Wiss. Berlin, 1869: 881. Possibly in Borborocoetes according to Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 257.

Borborocoetes pliciferus Werner, 1896, Verh. Zool. Bot. Ges. Wien, 46: 360. Holotype: ZMB; but MNHNP 1898.338 considered a syntype, apparently in error, by Ortiz and Lescure, 1990 "1989", Bull. Mus. Natl. Hist. Nat. Paris, Sect. A, Zool., 11: 115. Type locality: "Frutillar in Chile". Synonymy without discussion with Alsodes nodosus by Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 297. Synonymy with Eupsophus roseus by Ortiz and Lescure, 1990 "1989", Bull. Mus. Natl. Hist. Nat. Paris, Sect. A, Zool., 11: 115 (who rejected Cei's synonymy explicitly).

Cystignathus (Borborocoetus) grayi — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 84.

Cystignathus (Borborocoetus) bibronii — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 86.

Cystignathus (Borborocoetus) nigrita Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 88. Syntypes: MNHNC, according to the original publication. Type locality: "Habitat prope oppidum Valdivia,.... in praedio meo S. Jaun inveni", Valdivia, Chile. Synonymy (with Eupsophus grayi) by Cei, 1958, Invest. Zool. Chilen., 4: 275. Synonymy by Gorham, 1966, Das Tierreich, 85: 118; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Borborocoetus (Cystignathus) insularis var. ruber Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 91. Types: MNHNC, according to the original publication. Type locality: "Valdivia", Chile. Synonymy (with Eupsophus grayi) by Cei, 1958, Invest. Zool. Chilen., 4: 275. Synonymy by Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84. Previously placed by Gorham, 1966, Das Tierreich, 85: 118, as a synonym of Eupsophus roseus.

Borborocoetus fasciatus — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 92.

Cystignathus pallidus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 104. Syntypes: MNHNC (6 specimens), according to the original publication. Type locality: "prov. Valdivia", Chile. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Gorham, 1966, Das Tierreich, 85: 118; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Cystignathus (Borborocoetus) tympanicus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 106. Syntypes: MNHNC (2 specimens), according to the original publication. Type locality: not designated (presumably Chile). Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Gorham, 1966, Das Tierreich, 85: 118; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Cystignathus Boedeckeri Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 109. Types: MNHNC, according to the original publication. Type locality: "Concepcion", Chile. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84. Previously placed by Gorham, 1966, Das Tierreich, 85: 118, as a synonym of Eupsophus roseus.

Cystignathus oxyglossus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 110. Syntypes: MNHNC (7 specimens), according to the original publication. Type locality: "parte orientali insulae Chiloe", Chile. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Cystignathus fernandezi Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 112. Types: MNHNC, according to the original publication. Type locality: "insula Chiloé", Chile. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Cystignathus oreophilus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 117. Syntypes: MNHNC according to the original publication. Type locality: "cordillera de Cautin", Chile. Considered a nomen dubium by Cei, 1958, Invest. Zool. Chilen., 4: 276. Synonymy by Gorham, 1966, Das Tierreich, 85: 118; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Borborocoetes fasciatus — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 157.

Borborocoetes bibroni — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 157.

Borborocoetes nigrita — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 157.

Borborocoetes insularis var. ruber — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 157.

Cystignathus bödeckeri — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 158. Variant spelling.

Cystignathus oxiglossus — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Spelling error.

Leptodactylus kreffti Werner, 1904, Ergebn. Hamburger Magalhaens. Sammelr. 1892/93: 18. Holotype: Presumably ZMH although not stated. Type locality: "Coronel, in einem Wiesenteich", Chile. Considered by Heyer, 1974, Contrib. Sci. Nat. Hist. Mus. Los Angeles Co., 253: 43, to likely be referable to Eupsophus. Placed in the synonymy of Eupsophus roseus, without comment, by Gorham, 1966, Das Tierreich, 85: 118; Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Borborocoetes masareyi Roux, 1910, Zool. Anz., 36: 111. Holotype: NHMB 2789 according to Forcart, 1946, Verh. Naturforsch. Ges. Basel, 57: 122. Type locality: "Coronel, Chile". Synonymy (with Eupsophus grayi) by Cei, 1960, Breviora, 118: 1-13, and Cei, 1962, Invest. Zool. Chilen., 8: 33. Synonymy by Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 128, Gorham, 1966, Das Tierreich, 85: 118; and Lavilla, 1994 "1992", Acta Zool. Lilloana, 42: 84.

Leptodactylus oxyglossus — Nieden, 1923, Das Tierreich, 46: 492.

Leptodactylus boedeckeri — Nieden, 1923, Das Tierreich, 46: 492.

Leptodactylus oreophilus — Nieden, 1923, Das Tierreich, 46: 492.

Borborocoetes nigrita — Nieden, 1923, Das Tierreich, 46: 397.

Leptodactylus fernandezi — Nieden, 1923, Das Tierreich, 46: 492.

Leptodactylus pallidus — Nieden, 1923, Das Tierreich, 46: 492.

Platymantis kreffti — Lutz, 1930, Mem. Oswaldo Cruz, Rio de Janeiro, 23: 26.

Eupsophus roseus — Schmidt, 1954, Lunds Univ. Ärsskrift, N.F., Avd. 2, 49: 5; Cei, 1958, Invest. Zool. Chilen., 4: 275; Capurro-S., 1958, Invest. Zool. Chilen., 4: 294; Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 111–149.

Eupsophus bibroni — Capurro-S., 1958, Invest. Zool. Chilen., 4: 294.

Eupsophus grayi — Cei, 1958, Invest. Zool. Chilen., 4: 275; Capurro-S., 1958, Invest. Zool. Chilen., 4: 293.

Eupsophus grayii grayii — Capurro-S., 1963, Invest. Zool. Chilen., 10: 10-18, by implication.

Eupsophus sylvestris — Gorham, 1966, Das Tierreich, 85: 119.

Eupsophus roseus — Lynch, 1972, Bull. S. California Acad. Sci., 71: 7.

Common Names

Rosy Ground Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 81).

Distribution

Range poorly understood at present although as currently understood the northern limit in Chile is Tomé, Bíobio Region, and the southern limit is in the western foothills of the Andes at Los Mañíos (ca. 40° 20′ S); in Argentina, adjacent to Chile, it is found at Lago Paimún, at about 39°50′S, and Termas de Epulafquén, both in the province of Neuquen, although it may extend farther north in that country.

Geographic Occurrence

Natural Resident: Argentina, Chile

Comment

See account by Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 282–283. Formas and Vera, 1980, J. Herpetol., 14: 11–14, reported on advertisement call, larval morphology, and reproductive behavior and ecology. Penna and Veloso, 1990, J. Herpetol., 24: 23–33, reported on advertisement call. In the Eupsophus roseus group of Formas and Brieva, 1994, Proc. Biol. Soc. Washington, 107: 391–397. Márquez, Penna, Marques, and Amaral, 2005, Herpetol. J., 15: 257–263, reported on advertisement call. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 622, who noted (personal comm. from Esteban Lavilla) that records of this species from Argentina actually apply to Eupsophus calcaratus. See photograph, map, description of geographic range and habitat (as Eupsophus nahuelbuensis), and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 393. Revised by Correa-Quezada, Vásquez, Castro-Carrasco, Zúñiga-Reinoso, Ortiz, and Palma, 2017, PLoS One, 12(8: e0181026): 1–21, and Suárez-Villota, Quercia, Vera-Sovier, and Nuñez, 2018, PLoS One, 13(12: e0204968): 1–19 (who noted an unnamed cryptic species). See Lobos, Vidal-Maldonado, Correa-Quezada, Labra-Lillo, Díaz-Páez, Charrier, Rabanal, Díaz-Vidal, and Tala, 2013, Anf. Chile Conserv. 1–104, for photograph, comments on conservation status, and range. See Correa-Quezada and Durán, 2019, ZooKeys, 863: 107–152, for detailed discussion of systematics and range. See Charrier, 2019, Anf. Bosques Centro Sur y Patagonia de Chile: 194–197, for account (morphology, habitat, natural history) and polygon distribution map.

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