Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eupsophus vertebralis Grandison, 1961

Class: Amphibia > Order: Anura > Family: Alsodidae > Genus: Eupsophus > Species: Eupsophus vertebralis

Eupsophus vertebralis Grandison, 1961, Bull. Brit. Mus. (Nat. Hist.), Zool., 8: 136. Holotype: NHMW 4660.1, according to the original publication. Type locality: "Valdivia, Chile".

Common Names

Valdivia Ground Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 81).

Distribution

From the National Park Tolhuaca (Región de La Araucanía) to the Cordillera Pelada (Región de Los Ríos), Chile; reported from Puerto Blest, Rio Negro Province, Argentina, 50–1000 m elevation.

Geographic Occurrence

Natural Resident: Argentina, Chile

Comment

Considered a synonym of Alsodes vittatus by Donoso-Barros, 1976, Arch. Biol. Med. Experim., Univ. Chile, 10: 50. Removed from synonymy by Formas, 1989, Bol. Soc. Biol. Concepción, 60: 123–127. Penna and Veloso, 1990, J. Herpetol., 24: 23–33, reported on advertisement call. In the Eupsophus vertebralis group of Formas and Brieva, 1994, Proc. Biol. Soc. Washington, 107: 391–397. Formas, 1993, J. Herpetol., 28: 394–395, reported on a triploid individual. Basso and Úbeda, 1999, Herpetol. Rev., 30: 172, provided a record from Puerto Blest, Nahuel Huapi National Park, Bariloch Department, Río Negro, Argentina. See statement of geographic range, habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 622. See Lobos, Vidal-Maldonado, Correa-Quezada, Labra-Lillo, Díaz-Páez, Charrier, Rabanal, Díaz-Vidal, and Tala, 2013, Anf. Chile Conserv. 1–104, for photograph, comments on conservation status, and range. Suárez-Villota, Quercia, and Nuñez, 2018, J. Genomics, 6: 98–102, reported on the mitochondrial genome. See Correa-Quezada and Durán, 2019, ZooKeys, 863: 107–152, for discussion of systematics and range. See Charrier, 2019, Anf. Bosques Centro Sur y Patagonia de Chile: 202–205, for account (morphology, habitat, natural history) and polygon distribution map. Quercia, Suárez-Villota, Foresti, and Nuñez, 2020, Comp. Cytogenet., 14: 61–74, discussed the comparative cytogenetics of this species and Eupsophus emiliopugini.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.