Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Batrachyla leptopus Bell, 1843

Class: Amphibia > Order: Anura > Family: Batrachylidae > Genus: Batrachyla > Species: Batrachyla leptopus

Batrachyla leptopus Bell, 1843, Zool. Voyage Beagle, Part 5: 43. Holotype: BMNH 1947.2.16.96 (formerly 1845.5.25.29), according to A.G.C. Grandison in Cannatella, 1985, in Frost (ed.), Amph. Species World: 262. Type locality: "Valdivia", Chile

Batrachyla leptopodus Guichenot, 1848, in Gay (ed.), Hist. Fis. Polit. Chile, Zool., 2: 114. Error for Batrachyla leptopus Bell, 1843.

Hylodes (Batrachyla) leptopus — Cope, 1862, Proc. Acad. Nat. Sci. Philadelphia, 14: 153.

Hylodes leptopus — Cunningham, 1871, Trans. Linn. Soc. London, 27: 468; Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 93.

Hilodes chonotica Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 54. Types: MNHNC, according to the original publication. Type locality: "insulis Chonos", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Misspelling of generic name. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hylodes melanogastra Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 56. Holotype: MNHNC, according to the original publication. Type locality: "parte orientali insulae Chiloe", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hylodes brevipes Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 58. Holotype: MNHNC according to the original publication. Type locality: "oriental insulae Chiloé", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275. Provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hylodes macromeros Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 59. Syntypes: MNHNC (3 specimens), according to the original publication. Type locality: unknown (presumably Chile). Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Provisional synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; Gorham, 1966, Das Tierreich, 85: 34.

Hylodes fitzingeri Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 62. Syntypes: MNHNC (2 specimens), according to the original publication. Type locality: "privincia Valdivia ad flumen Pelmaiquén", Chile. Preoccupied by Hylodes fitzingeri Schmidt, 1857. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34. Junior homonym of Hylodes fitzingeri Schmidt, 1857 (= Craugastor fitzingeri).

Hylodes stenocephala Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 63. Syntypes: MNHNC (2 specimens), according to the original publication. Type locality: "Concepcion", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275. Provisional synonym of Gorham, 1966, Das Tierreich, 85: 34.

Hylodes cardioglossa Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 64. Holotype: MNHNC, according to the original publication. Type locality: "insula Chiloé", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hylodes granulatus Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 66. Syntypes: MNHNC (4 specimens), according to the original publication. Type locality: "parte insulae Chiloé", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hylodes gracilis Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 67. Types: MNHNC (6 specimens), according to the original publicationi. Type locality: "provincia Valdivia", Chile. Considered a nomen dubium by Nieden, 1923, Das Tierreich, 46: 467. Synonymy by Cei, 1958, Invest. Zool. Chilen., 4: 275; provisional synonymy by Gorham, 1966, Das Tierreich, 85: 34.

Hilodes brevipes — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect subsequent spelling of generic name.

Hilodes cardioglossa — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect subsequent spelling.

Hilodes fitzingeri — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect spelling of the generic name.

Hilodes granulatus — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect subsequent spelling of generic name.

Hilodes leptopus — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect spelling of generic name.

Hilodes macromeros — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect spelling of generic name.

Hilodes melanogastra — Philippi, 1902, Supl. Batr. Chil. Descr. Hist. Fis. Polit. Chile: 159. Incorrect spelling of generic name.

Eleutherodactylus leptopus — Myers, 1962, Copeia, 1962: 197, by implication.

Common Names

Gray Wood Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 70).

Distribution

From Los Queules (Región del Maule) to the Río Baker (Región de Aysén), Chile; in Nothofagus forests from Neuquén to lower Río Negro, Argentina.

Geographic Occurrence

Natural Resident: Argentina, Chile

Comment

See accounts by Cei, 1962, Batr. Chile: 83–85, and Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 275–277. Penna and Veloso, 1990, J. Herpetol., 24: 23–33, reported on advertisement call. Díaz-Páez and Ortiz, 2006, Herpetol. Rev., 37: 485-486, provided specific localities and a range extension. Vidal-Maldonado and Ortiz, 2009, Check List, 5: 869–871, reported the species from Butachauques and Talcán Islands, between Chiloé and the mainland, Chile, and discussed the biogeography of the species. Cuevas and Cifuentes P., 2010, Check List, 6: 633–636, provided a northern-most record and disucssed and mapped the range. Blotto, Nuñez, Úbeda, Wheeler, and Faivovich, 2013, Cladistics, 29: 113–131, regarded this species to be in Batrachyla on the basis of molecular phylogenetics. See Lobos, Vidal-Maldonado, Correa-Quezada, Labra-Lillo, Díaz-Páez, Charrier, Rabanal, Díaz-Vidal, and Tala, 2013, Anf. Chile Conserv. 1–104, for photograph, comments on conservation status, and range. See Charrier, 2019, Anf. Bosques Centro Sur y Patagonia de Chile: 146–149, for account (morphology, habitat, natural history) and polygon distribution map.

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