Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Atopophrynus Lynch and Ruiz-Carranza, 1982

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Genus: Atopophrynus

1 species

Atopophrynus Lynch and Ruiz-Carranza, 1982, Proc. Biol. Soc. Washington, 95: 557. Type species: Atopophrynus syntomopus Lynch and Ruiz-Carranza, 1982, by original designation.

English Names

Sonson Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 70).

Distribution

As for the single species.

Geographics occurrence

Natural resident: Colombia

Endemic to the political unit: Colombia

Comment

Considered a member of Dendrobatidae in the original publication. Placed in Leptodactylidae (sensu lato), possibly close to Geobatrachus, by Myers and Ford, 1986, Am. Mus. Novit., 2843: 1-15, although they also noted that it might fit into Bufonidae. Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, 104: 10092-10097, did not address this taxon in their analysis so its phylogenetic placement is unknown. Assignment to their Strabomantidae or Strabomantinae by Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 1-182, was provisional. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, also did not include this taxon in their analysis and placed it provisionally as incertae sedis within Craugastoridae. Padial, Grant, and Frost, 2014, Zootaxa, 3825: 122, transferred Atopophrynus and Geobatrachus to incertae sedis within their Brachycephaloidea (= Terrarana of other authors), but did not assign these genera to families within that taxon pending resolution of their phylogenetic position.

Contained taxa (1 sp.):

Atopophrynus syntomopus Lynch and Ruiz-Carranza, 1982

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