Ceuthomantidae Heinicke, Duellman, Trueb, Means, MacCulloch, and Hedges, 2009

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Ceuthomantidae
4 species

Ceuthomantidae Heinicke, Duellman, Trueb, Means, MacCulloch, and Hedges, 2009, Zootaxa, 2211: 6. Type genus: Ceuthomantis Heinicke, Duellman, Trueb, Means, MacCulloch, and Hedges, 2009.

Ceuthomantinae — Padial, Grant, and Frost, 2014, Zootaxa, 3827: 599.

English Names

Emerald-barred Frogs (Vitt and Caldwell, 2014, Herpetology, 4th Ed.: 498).


Elevations of 493–1540 m in the southern and eastern parts of the Guiana Highlands, including the slopes and tops of the following: Mount Ayanganna and the Wokomung Massif in Guyana, Cerro Aracamuni and Sierra Tapirapecó in the Cerro Neblina Massif on the Venezuela–Brazil border, and Sarisariñama Tepui in southern Venezuela, extending into adjacent Brazil.


Ceuthomantidae (in the sense of only including Ceuthomantis) was considered the sister taxon of the remaining members of Terrarana/Brachycephaloidea (Craugastoridae and Eleutherodactylidae of this catalogue) according to the original publication. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543–583, confirmed this relationship using similar inferential methods and a largely similar dataset. Padial, Grant, and Frost, 2014, Zootaxa, 3825: 125, discovered that that this topology was highly alignment and tree-building method dependent and transferred Ceuthomantis into Pristimantinae, for which the name Ceuthomantinae is the oldest name. Blackburn and Wake, 2011, In Zhang (ed.), Zootaxa, 3148: 39–55, briefly reviewed the nomenclatural history of this taxon. Jetz and Pyron, 2018, Nature Ecol. & Evol., 2: 850–858, Heinicke, Lemmon, Lemmon, McGrath, and Hedges, 2017 "2018", Mol. Phylogenet. Evol., 118: 145–155, and Motta, Taucce, Haddad, and Canedo, 2021, J. Zool. Syst. Evol. Res., 59: 663–679, provided further evidence for Ceuthomantidae being monogeneric and the sister taxon of the remaining brachycephaloids. Cole, Townsend, Reynolds, MacCulloch, and Lathrop, 2013, Proc. Biol. Soc. Washington, 125: 317–578, provided identification keys and accounts for the species in Guyana. Ospina-Sarria and Grant, 2022, Zool. J. Linn. Soc., 195: 976–994, reported on morphology of pelivic and thigh musculature and digital discs of the Brachycephaloidea, and identified an unambiguous synapomorphy of Ceuthomantidae.  

Contained taxa (4 sp.):

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