Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eleutherodactylus atkinsi Dunn, 1925

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Eleutherodactylidae > Subfamily: Eleutherodactylinae > Genus: Eleutherodactylus > Species: Eleutherodactylus atkinsi

Eleutherodactylus atkinsi Dunn, 1925, Occas. Pap. Boston Soc. Nat. Hist., 5: 165. Holotype: MCZ 10587, by original designation. Type locality: "Colonia Guabairo, near Cienfuegos, [Cienfuegos Province,] Cuba". Rendered as "Near Soledad, Cienfuegos, Cuba" by Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 256.

Eleutherodactylus atkinsi orientalis Barbour and Shreve, 1937, Bull. Mus. Comp. Zool., 80: 392. Holotype: MCZ 22158, by original designation. Type locality: "Upper Ovando River, 1000–1200 feet, Oriente [Province; now Guantánamo Province], Cuba".

Eleutherodactylus atkinsi atkinsi — Barbour and Shreve, 1937, Bull. Mus. Comp. Zool., 80: 392.

Eleutherodactylus (Euhyas) atkinsi — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 325; Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.

Eleutherodactylus (Euhyas) atkinsi estradai — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 325, by implication.

Eleutherodactylus atkinsi estradai Lynch, 1991, Copeia, 1991: 1138. Replacement name for Eleutherodactylus atkinsi orientalis Barbour and Shreve, 1937, which is a preoccupied secondary homonym of Sminthillus limbatus orientalis Barbour and Shreve, 1937 (= Eleutherodactylus orientalis).

Eleutherodactylus (Euhyas) estradai — Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 223. Unintended combination.

Euhyas atkinsi — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 361; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 433.

Common Names

Atkins' Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 72).

Cuban Groin-spotted Frog (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 9).

Distribution

Widespread in Cuba and the Isla de Juventud (Isla de Pinos), sea leve to 1212 m elevation.

Geographic Occurrence

Natural Resident: Cuba

Endemic: Cuba

Comment

In the Eleutherodactylus ricordii group, according to Schwartz, 1958, Am. Mus. Novit., 1873: 2; two subspecies recognized. Not assignable to species group according to Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 325. In the Eleutherodactylus (Euhyas) ricordii group according to Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 220. See Eleutherodactylus estradai. Díaz and Cádiz, 2007, Herpetotropicos, Mérida, 3: 100–122, reported on the advertisement call. In the Eleutherodactylus (Euhyas) planirostris species series, monotypic Eleutherodactylus atkinsi species group of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 78, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 131. Díaz and Cádiz, 2008, Guía Taxon. Anf. Cuba: 82–83, provided a brief account, illustration, and map. Henderson and Powell, 2009, Nat. Hist. Rept. Amph. W. Indies: 37, summarized the natural history literature. Rivalta González, Rodríguez Schettino, Mancina, and Iturriaga, 2014, Smithson. Herpetol. Inform. Serv., 145: 22–23, provided a dot map and localities. Rodriguez, Börner, Pabijan, Gehara, Haddad, and Vences, 2015, Evol. Ecol., 29: 765–785 (and supplemental data), reported on phylogeographic diversity and its causes.

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