Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eleutherodactylus varians (Gundlach and Peters, 1864)

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Eleutherodactylidae > Subfamily: Eleutherodactylinae > Genus: Eleutherodactylus > Species: Eleutherodactylus varians

Hylodes varians Gundlach and Peters In Peters, 1864, Monatsber. Preuss. Akad. Wiss. Berlin, 1864: 390. Syntypes: ZMB 5108, MCZ 11621 (on exchange from ZMB, according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 281) according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 46. Type locality: "Cuba".

Eleutherodactylus varians — Stejneger, 1904, Annu. Rep. U.S. Natl. Mus. for 1902: 582-583, by implication; Barbour, 1914, Mem. Mus. Comp. Zool., 44: 245.

Eleutherodactylus varians varians — Schwartz, 1960, Sci. Publ. Reading Public Mus. Art Gallery, 11: 8.

Eleutherodactylus (Eleutherodactylus) varians — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 329; Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: 10.

Common Names

Cliff Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 80).

Cuban Bromeliad Frog (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 12).

Distribution

Central and eastern Cuba from extreme northeastern Artemesia Province east in increasingly isolated population to Guantanamo and Granma Provinces, sea level to 845 m elevation.

Geographic Occurrence

Natural Resident: Cuba

Endemic: Cuba

Comment

I In the Eleutherodactylus auriculatus group, according to Schwartz, 1958, Am. Mus. Novit., 1873: 2; and Schwartz, 1969, Stud. Fauna Curaçao and other Caribb. Is., 30: 100In the Eleutherodactylus auriculatus section, unassigned to series or species group, according to Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 329. Hedges, Estrada, and Thomas, 1992, Herpetol. Monogr., 6: 68–83, reported on the call. Díaz and Cádiz, 2007, Herpetotropicos, Mérida, 3: 100–122, reported on the advertisement call. In the Eleutherodactylus (Eleutherodactylus) martinicensis series, Eleutherodactylus martinicensis group according to Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 229. In the Eleutherodactylus (Eleutherodactylus) varians species series, Eleutherodactylus varians species group, Eleutherodactylus varians species subgroup of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 62–64, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 130. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 388. Díaz and Cádiz, 2008, Guía Taxon. Anf. Cuba: 120–121, provided a brief account, illustration, and map (in the sense of including Eleutherodactylus olibrus and Eleutherodactylus staurometopon as subspecies). Henderson and Powell, 2009, Nat. Hist. Rept. Amph. W. Indies: 80–81, summarized the natural history literature. Rivalta González, Rodríguez Schettino, Mancina, and Iturriaga, 2014, Smithson. Herpetol. Inform. Serv., 145: 16, provided a dot map and localities.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.