Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Eleutherodactylus varleyi Dunn, 1925

Class: Amphibia > Order: Anura > Superfamily: Brachycephaloidea > Family: Eleutherodactylidae > Subfamily: Eleutherodactylinae > Genus: Eleutherodactylus > Species: Eleutherodactylus varleyi

Eleutherodactylus varleyi Dunn, 1925, Occas. Pap. Boston Soc. Nat. Hist., 5: 163. Holotype: MCZ 10601, by original designation. Type locality: "Soledad, [Cienfuegos Province,] Cuba".

Eleutherodactylus phyzelus Schwartz, 1958, Am. Mus. Novit., 1873: 7. Holotype: AMNH 59832, by original designation. Type locality: "4.4 miles northwest of San Vicente, on road between San Vicente and Puerto Esperanza, Pinar del Río Province, Cuba". Synonymy by Schwartz, 1964, Breviora, 208: 9.

Eleutherodactylus (Euhyas) varleyi — Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 325; Heinicke, Duellman, and Hedges, 2007, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 104: Table 2.

Euhyas varleyi — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 361; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 433.

English Names

Varley's Robber Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 80).

Western Cuba Grassfrog (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 12).

Distribution

Western Sancti Spiritus Province and to the west in Cuba as well as the Isla de Juventud, sea level to 900 m elevation.

Geographic Occurrence

Natural Resident: Cuba

Endemic: Cuba

Comment

In the Eleutherodactylus varleyi group, according to Schwartz, 1958, Am. Mus. Novit., 1873: 2; Shreve and Williams, 1963, Bull. Mus. Comp. Zool., 129: 339; and Hedges, 1989, in Woods (ed.), Biogeograph. W. Indies: 325. In the Eleutherodactylus (Euhyas) unassigned to species group according to Lynch and Duellman, 1997, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 23: 234. Díaz and Cádiz, 2007, Herpetotropicos, Mérida, 3: 100–122, reported on the advertisement call. In the Eleutherodactylus (Euhyas) planirostris species series, Eleutherodactylus gundlachi species group of Hedges, Duellman, and Heinicke, 2008, Zootaxa, 1737: 79-80, and of Padial, Grant, and Frost, 2014, Zootaxa, 3825: 131.Díaz and Cádiz, 2008, Guía Taxon. Anf. Cuba: 68-69, provided a brief account, illustration, and map. Díaz, Hedges, and Schmid, 2012, Zootaxa, 3220: 44-60, redelimited the species by partitioning out Eleutherodactylus feichtingeri and noted another cryptic species hiding under this name in Artemesia Province, Cuba. Henderson and Powell, 2009, Nat. Hist. Rept. Amph. W. Indies: 81, summarized the natural history literature.

External links:

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For information on distribution, habitat, and conservation see the Map of Life
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For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.