Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Atelopus ebenoides Rivero, 1963

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Atelopus > Species: Atelopus ebenoides

Atelopus ebenoides Rivero, 1963, Caribb. J. Sci., 3: 120. Holotype: FMNH 69746, by original designation. Type locality: "Páramo de las Papas, 3,600 m., San Agustín, Huila, Colombia".

Atelopus ebenoides ebenoides — Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 122.

Atelopus eusebianus Rivero and Granados-Díaz, 1993, Caribb. J. Sci., 29: 12. Holotype: ICN 20010, by original designation. Type locality: "Malvasa, Municipio de Totoró, 2° 3′ N, 76° 24′ W, 2995 m, Depto. de Cauca, Colombia". Synonymy by Plewnia, Dias, Anganoy-Criollo, Venegas-Valencia, Szepanski, Heine, Böning, Ramírez, and Lötters, 2025, Phyllomedusa, 24: 223.

Atelopus angelito Ardila-Robayo and Ruiz-Carranza, 1998, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 22: 281. Holotype: ICN 33408, by original designation. Type locality: "COLOMBIA, Departamento del Cauca, Municipio de San Sebastián, Inspección de policía Valencia, ca. Km. 5-6 por camino de herradura de Valencia a Quinchana, ca. 1° 51′ latitud N, 76° 47′ longitud W de Greenwich, 2900 m-3000m". Synonymy by Plewnia, Dias, Anganoy-Criollo, Venegas-Valencia, Szepanski, Heine, Böning, Ramírez, and Lötters, 2025, Phyllomedusa, 24: 223.

Common Names

Huila Stubfoot Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 39).

Angelito's Harlequin Frog (Atelopus angelito [no longer recognized]: Coloma and Duellman, 2025, Amph. Ecuador. Vol. 2: xxx).

Rana Arlequín de Angelito (Atelopus angelito [no longer recognized]: Spanish: Coloma and Duellman, 2025, Amph. Ecuador. Vol. 2: xxx).

Malvasa Stubfoot Toad (Atelopus eusebianus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 39).

Distribution

Southern part of the central Andes (eastern Cauca, southwestern Huila, and southern Tolima departments) of Colombia, in paramo and subparamo habitats, 2500–3700 m elevation.

angelito: Elevations of 2500 to 3000 m along the eastern slope of the Cordillera Occidental from the Departamento Valle del Cauca in Colombia to the Cordillera Central in Carchi Province, Ecuador.

eusebianus: Cloud forest on the western slope of the Central Andes in the Departments of Cauca and Valle del Cauca (Pradera municipality; see comment), Colombia, 1050–3250 m elevation.

Geographic Occurrence

Natural Resident: Colombia, Ecuador

Comment

In the Atelopus ignescens group of Lynch, 1993, Alytes, 11: 77–87. See Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 122–125, for subspecies and account. P. Ruiz, in Lötters, 1996, Neotrop. Toad Genus Atelopus: 24, suggested that the population at the Tolima locality represents an unnamed species. Bolívar-García and Restrepo-Toro, 2004, in Rueda-Almonacid et al. (eds.), Libro Rojo Anf. Colombia: 185–188, provided an account (for nominal Atelopus eusebianus) and map. Gutiérrez and López-López, 2005, in Rueda-Almonacid et al. (eds.), Ranas Arlequines: 70, provided a brief account, photograph, and map. Ardila-Robayo, 2005, in Rueda-Almonacid et al. (eds.), Ranas Arlequines: 56, provided a brief account for nominal Atelopus angelito. Valencia-Rojas and López-López, 2005, Nov. Colomb., N. Epoca, 8: 71–74, reported an additional locality for Atelopus eusebianus. López-López and Granados-Díaz, 2005, in Rueda-Almonacid et al. (eds.), Ranas Arlequines: 74, provided a brief account, photograph, and map for nominal Atelopus eusebianus. See photograph, map, description of geographic range and habitat, and conservation status of Atelopus ebenoides in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 160–165 (in the sense of including Atelopus marinkellei); they also provided similar accounts for Atelopus eusebianus and Atelopus angelito. Coloma, Duellman, Almendáriz C., Ron, Terán-Valdez, and Guayasamin, 2010, Zootaxa, 2574: 1–54, discussed the Ecuador record of Atelopus angelito and noted that the population may be extinct. Hernández-Córdoba, Cardona-Botero, and Castro-Herrera, 2014, Check List, 10: 682–683, provided a record of nominal Atelopus eusebianus for Valle del Cauca, Colombia, and mapped the range of the species, although this identification requires genetic confirmation. Coloma and Duellman, 2025, Amph. Ecuador. Vol. 2: 201–202, provided an account of nominal Atelopus angelito, with photographs, which summarized identification, adult morphology, systematics, natural history, distribution (including a dot map for Ecuador), and conservation.

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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador