Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Atelopus harlequin Coloma, Plewnia, Böning, Boistel, Ellwein, Lötters, Paluh, Roca-Rey Ross and Venegas, 2025

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Atelopus > Species: Atelopus harlequin

Atelopus harlequin Coloma, Plewnia, Böning, Boistel, Ellwein, Lötters, Paluh, Roca-Rey Ross and Venegas in Lötters, Böning, Bailon, Barros Castañeda, Boistel, Catenazzi, Chaparro, Chávez, Chujutalli, Coen, Coloma, Crawford, Culebras, Cusi Martínez, Daza-R., De la Riva, Ellwein, Ernst, Flechas-Hernández, Fouquet, Guayasamin, Heine, Jorge, Jung, Jungfer, Kaffenberger, Krehenwinkel, La Marca, Lampo, Medina-Rangel, Orsen, Paluh, Peréz-González, Perrin, Quezada Riera, Reyes-Puig, Roca-Rey Ross, Rößler, Rueda-Solano, Salazar-Valenzuela, Señaris Vasquez, Sowinski, Terán-Valdez, Tovar-Ortiz, Veith, Venegas, von May, Weitkamp, and Plewnia, 2025, Zootaxa, 5571: 44. Holotype: MUBI 2448, by original designation. Type locality: "PERU: Loreto: km 74 on road from Iquitos to Nauta along a tributary of the Río Itaya (4°17’08”S, 73°31’45”W, ca. 100 m asl)". Zoobank Publication registration: AB79DE73-A1FD-40EB-8024-8CDB1DF1AE90

Common Names

None noted.

Distribution

Known from various localities between the Río Napo (to its right) and the Río Marañon (to its left) and west of the Amazon River at ca. 100–150 m elevation; geographic range is delimited by the latter two rivers, while the distributional limits to the North remain difficult to assess. In Ecuador, this species is known only from Lorocachi in Pastaza Province, situated in the upper Amazon basin at ca. 220 m asl, in the Río Pastaza drainage. Unknown if the area between Lorocachi and the northern and western banks of the Marañon and Amazon Rivers is continuously colonized as the area remains poorly studied and currently known only from two localities inbetween (Pucacuro and Andoas).

Geographic Occurrence

Natural Resident: Ecuador, Peru

Comment

Delimited on the basis of call, osteology, adult morphology, and molecular markers in the original publication from Atelopus spumarius where they also addressed confusion in previous literature.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador