Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Atelopus tricolor Boulenger, 1902

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Atelopus > Species: Atelopus tricolor

Atelopus tricolor Boulenger, 1902, Ann. Mag. Nat. Hist., Ser. 7, 10: 397. Syntypes: BMNH 1947.2.14.57–59 (formerly 1902.5.29.192–195), acccording to museum records, and ZFMK 28103 (according to Böhme and Bischoff, 1984, Bonn. Zool. Monogr., 19: 177), USNM 193575 (formerly BMNH 1947.2.14.60) (according to Peters, 1973, Smithson. Contrib. Zool., 145: 43); ZFMK 28103 designated lectotype by Lötters and De la Riva, 1998, J. Herpetol., 32: 484. Type locality: "Marcapata Valley, [Departamento Cusco,] E. Peru".

Atelopus rugulosus Noble, 1921, Am. Mus. Novit., 29: 3. Holotype: AMNH 6097, by original designation. Type locality: "vicinity of Juliaca, Peru". Type locality doubted by Lötters, 1996, Neotrop. Toad Genus Atelopus: 52. Synonymy by Reynolds and Foster, 1992, Herpetol. Monogr., 6: 86.

Atelopus willimani Donoso-Barros, 1969, Physis, Buenos Aires, 28: 327. Holotype: IZUC (formerly R. Donoso-Barros 660) according to Hoogmoed, 1985, in Frost (ed.), Amph. Species World: 34. Type locality: "Rio Beni, Runerrabaque [= Rurrenabaque], Bolivia". Synonymy by Lötters and De la Riva, 1998, J. Herpetol., 32: 481–488.

Common Names

Three-colored Stubfoot Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 40).

Three-colored Harlequin Toad (Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 177).

Bolivian Stubfoot Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 40 [Atelopus willimani—no longer recognized).

Distribution

Humid montane forests of the Cordillera Oriental of southeastern Peru to central Bolivia, between 1250–2500 m elevation.

Geographic Occurrence

Natural Resident: Bolivia, Peru

Comment

Likely in the Atelopus flavescens group of Lynch, 1993, Alytes, 11: 77–87. Larval morphology described by Lavilla, de Sá, and De la Riva, 1997, J. Herpetol., 31: 121–124. See accounts by Lötters and De la Riva, 1998, J. Herpetol., 32: 481–488, and Köhler, 2000, Bonn. Zool. Monogr., 48: 80–81. Lötters, Glaw, Reichle, Köhler, and Meyer, 1999, Herpetozoa, Wien, 12: 79–83, discussed vocalization. Lavilla and de Sá, 2001, Amphibia-Reptilia, 22: 167–177, reported on the larval chondrocranium and visceral skeleton. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 177. Chaparro Auza and Cortez-Fernandez, 2005, in Rueda-Almonacid et al. (eds.), Ranas Arlequines: 128, provided a brief account, photograph, and map. De la Riva and Reichle, 2014, Herpetol. Monogr., 28: 59, who discussed conservation status, noted that attempts to find extant populations in Bolivia have been unsuccessful and that the species was last seen in that country in December 2003. Pacheco-Suarez, 2020, Salamandra, 56: 392–394, discussed rediscovery of the species in Bolivia in 2020. Ocampo, Aparicio, Bernal Hoverud, Domic, and Wallace, 2024, Herpetol. Notes, 17: 371–389, reported the species and its habitat from Madidi National Park, La Paz, Bolivia.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist