Bufo tuberculatus Zarevskij, 1926

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufo > Species: Bufo tuberculatus

Bufo tuberculatus Zarevskij, 1926 "1925", Annu. Mus. Zool. Acad. Sci. Leningrad, 26: 75. Syntypes: ZISP 1969 (plateau of Kham), 1972 (plateau of Kham), 2634 (Mountains Ji-us-shan, lake Kuku-nor); ZISP 2634 designated lectotype by Borkin and Matsui, 1987 "1986", in Anajeva and Borkin (eds.), Sistematika i ekologiia amfibii i reptilii: 51; the identity of this lectotype questioned by Fei and Ye, 2016, Amph. China, 1: 761, who thought it might be a representative of Bufo gargarizans minshanicus. Type locality: "Tibet, plateau of Kham" (actually Qinghai Province, China), and "Mountains Ji-us-shan mountains, lake Kuku-nor [= Qinghai Lake]", Xizang, China.

Bufo (Bufo) tuberculatusDubois and Bour, 2010, Zootaxa, 2447: 24.

English Names

Qinghai Lake Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 43; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31).

Round-warted Toad (Fei, 1999, Atlas Amph. China: 138; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31).


Montane southwestern China in western Sichuan (Baitang, Xiangcheng), very northern Yunnan (Deqen), Qinghai, and the eastern tip of Xizang (Markam, 2600-2700 m elevation. See comment. 

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of


Resurrected from synonymy of Bufo tibetanus by Borkin and Matsui, 1987 "1986", in Anajeva and Borkin (eds.), Sistematika i ekologiia amfibii i reptilii: 43-54, where it had been placed by Liu, 1950, Fieldiana, Zool. Mem., 2: 207; in the synonymy of Bufo gargarizans by Pope and Boring, 1940, Peking Nat. Hist. Bull., 15: 32. Fei, 1999, Atlas Amph. China: 138-139, provided a brief account, figure, and map. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 159, provided a brief account and illustration. See statement of geographic range, habitat, and conservation status (as Bufo tuberculatus) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 612. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 546-550, provided an account for China and a spot map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 241, provided a brief account (as Bufo tuberculatus) including photographs of specimens and habitat. Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31, provided an account for Xizang, China. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 255, provided an account, photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 759–761, provided an account, photograph, and dot map. Dufresnes and Litvinchuk, 2022, Zool. J. Linn. Soc., 195: 700, considered this taxon to be a junior synonym of Bufo gargarizans on the basis of genetic distance. Retained here pending careful evaluation of the entire complex. 

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