Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Bufo tuberculatus Zarevskij, 1926

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufo > Species: Bufo tuberculatus

Bufo tuberculatus Zarevskij, 1926 "1925", Annu. Mus. Zool. Acad. Sci. Leningrad, 26: 75. Syntypes: ZISP 1969 (plateau of Kham), 1972 (plateau of Kham), 2634 (Mountains Ji-us-shan, lake Kuku-nor); ZISP 2634 designated lectotype by Borkin and Matsui, 1987 "1986", in Anajeva and Borkin (eds.), Sistematika i ekologiia amfibii i reptilii: 51; the identity of this lectotype questioned by Fei and Ye, 2016, Amph. China, 1: 761, who thought it might be a representative of Bufo gargarizans minshanicus. Type locality: "Tibet, plateau of Kham" (actually Qinghai Province, China), and "Mountains Ji-us-shan mountains, lake Kuku-nor [= Qinghai Lake]", Xizang, China.

Bufo (Bufo) tuberculatus — Dubois and Bour, 2010, Zootaxa, 2447: 24.

Common Names

Qinghai Lake Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 43; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31).

Round-warted Toad (Fei, 1999, Atlas Amph. China: 138; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31).

圆疣蟾蜍 (Chinese: Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang, and Che, 2020, Biodiversity Sci., 28: App. 1: 7).

Distribution

Montane southwestern China in western Sichuan (Baitang, Xiangcheng), very northern Yunnan (Deqen), Qinghai, and the eastern tip of Xizang (Markam, 2600-2700 m elevation. See comment.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Resurrected from synonymy of Bufo tibetanus by Borkin and Matsui, 1987 "1986", in Anajeva and Borkin (eds.), Sistematika i ekologiia amfibii i reptilii: 43-54, where it had been placed by Liu, 1950, Fieldiana, Zool. Mem., 2: 207; in the synonymy of Bufo gargarizans by Pope and Boring, 1940, Peking Nat. Hist. Bull., 15: 32. Fei, 1999, Atlas Amph. China: 138-139, provided a brief account, figure, and map. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 159, provided a brief account and illustration. See statement of geographic range, habitat, and conservation status (as Bufo tuberculatus) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 612. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 546-550, provided an account for China and a spot map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 241, provided a brief account (as Bufo tuberculatus) including photographs of specimens and habitat. Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 31, provided an account for Xizang, China. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 255, provided an account, photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 759–761, provided an account, photograph, and dot map. Dufresnes and Litvinchuk, 2022, Zool. J. Linn. Soc., 195: 700, considered this taxon to be a junior synonym of Bufo gargarizans on the basis of genetic distance. Fu, 2023, Asian Herpetol. Res., 14: 1–10, questioned the status of this taxon as seperable from Bufo gargarizans or Bufo tibetanus, citing specific issues of sampling (or lack thereof), conflicting evidence, and apparent misapplication of names, but made no taxonomic changes. Retained here pending careful evaluation of the entire complex.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China