Bufotes baturae (Stöck, Schmid, Steinlein, and Grosse, 1999)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Bufotes > Species: Bufotes baturae

Bufo pseudoraddei baturae Stöck, Schmid, Steinlein, and Grosse, 1999, Ital. J. Zool., 66: 221. Holotype: ZSM 103/1998, by original designation. Type locality: "a plain above the right bank of the Hunza river near the mouth of the Shimsal river, north of the village of Pasu, 2700 m a.s.l., Karakoram, Pakistan". Considered a triploid taxon by Stöck, Frynta, Grosse, Steinlein, and Schmid, 2001, Asiat. Herpetol. Res., 9: 81.

Bufo (Bufotes) pseudoraddei baturae — Dubois and Bour, 2010, Zootaxa, 2447: 25. 

Bufotes pseudoraddei baturae — Frost, 2013, Amph. Spec. World, Vers. 5.6. Required change because of non-monophyly of Bufo.

Bufo baturuaeBetto-Colliard, Sermier, Litvinchuk, Perrin, and Stöck, 2015, Heredity, 114: 301. 

Bufotes baturae — Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 17. 

English Names

Batura Toad (Bufo pseudoraddei baturaeStöck, Schmid, Steinlein, and Grosse, 1999, Ital. J. Zool., 66: 221).

Batura Glacier Toad (Bufo pseudoraddei baturaeKhan, 2002, Bull. Chicago Herpetol. Soc., 37: 159).

Distribution

Extreme northern Pakistan through northeastern Afghanistan to southeastern Tajikistan. 

Comment

A triploid lineage considered a distinct species of hybrid origin by Betto-Colliard, Sermier, Litvinchuk, Perrin, and Stöck, 2015, Heredity, 114: 301. Stöck, Lamatsch, Steinlein, Epplen, Grosse, Hock, Klapperstück, Lampert, Scheer, Schmid, and Schartl, 2002, Nature Genetics, 30: 325-328, provided cytogenetic details for Bufo pseudoraddei baturae. See comment under Bufotes pseudoraddei for additional relevant literature. See localities in Afghanistan and comments by Wagner, Bauer, Leviton, Wilms, and Böhme, 2016, Proc. California Acad. Sci., Ser. 4, 63: 463–464. See discussion and map by Ficetola and Stöck, 2016, J. Biogeograph., 43: 703–715. A triploid hybrid species (Bufotes turanensis X Bufotes latastii) according to Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 17. : 463. A triploid hybrid species (♀ Bufotes turanensis × ♂ Bufotes latastii) according to Dufresnes, Mazepa, Jablonski, Oliveira, Wenseleers, Shabanov, Auer, Ernst, Koch, Ramírez-Chaves, Mulder, Simonovo, Tiutenko, Kryvokhyzhar, Wennekes, Zinenko, Korshunov, Al-Johany, Peregontsev, Masroor, Betto-Colliard, Denoël, Borkin, Skorinov, Pasynkova, Mazanaeva, Rosanov, Dubey, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 141: 1–25, who summarized its biogeography and evolutionary history. Rais, Ahmed, Sajjad, Akram, Saeed, Hamid, and Abid, 2021, ZooKeys, 1062: 157–175, included this species in an identification key to the amphibian species of Pakistan as well as providing a photograph. 

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