Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ingerophrynus claviger (Peters, 1863)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Ingerophrynus > Species: Ingerophrynus claviger

Bufo claviger Peters, 1863, Monatsber. Preuss. Akad. Wiss. Berlin, 1863: 405. Syntypes: ZMB 37322 and 4863 according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 40, and (according to Marx, 1958, Fieldiana, Zool., 36: 427) FMNH 73840; Marx, 1958, Fieldiana, Zool., 36: 427, mentioned a "lectotype in ZMB" but did not cite a specimen; ZMB 4863 designated lectotype by Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 40. See also Bauer, 2016, in Das and Tuen (eds.), Nat. Explor. Field Sci. SE Asia Australasia: 93. Type locality: "Benkulen [= Bengkulu] auf Sumatra", Indonesia.

Ingerophrynus claviger — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365.

English Names

Benkulen Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 41).

Distribution

Nias I. and northern Bengkulu Province at low elevations, Sumatra, Indonesia.

Comment

Related to Ingerophrynus biporcatus (as Bufo) according to Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 312. In the former Bufo biporcatus group of Inger, 1972, in Blair (ed.), Evol. Genus Bufo: 107. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 26, to compare this species with others in the Sunda Shelf region (as Bufo). See map, description of geographic range and habitat, and conservation status (as Bufo claviger) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 181, who questioned the Nias I. record.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.