Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhaebo ceratophrys (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhaebo > Species: Rhaebo ceratophrys

Bufo ceratophrys Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 319, pl. 22, fig. 2. Holotype: BMNH 1880.12.5.151, according to Hoogmoed, 1985, in Frost (ed.), Amph. Species World: 40. Type locality: "Ecuador".

Rhinella ceratophrys — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 365.

Rhaebo ceratophrys — Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 37.

English Names

Horned Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 41).

Distribution

Upper Amazonian Colombia, southern Venezuela (Cerro Neblina and Cerro Marahuaca), eastern Ecuador, northeastern Peru (Loreto), and intervening border areas of western Amazonian Brazil.

Comment

See Hoogmoed, 1977, Zool. Meded., Leiden, 51: 265–275, and Hoogmoed, 1990, in Peters and Hutterer (eds.), Vert. Tropics: 113–123, for discussion. Rodríguez and Duellman, 1994, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 22: 12–13, provided a brief account. Barrio-Amorós, 1999 "1998", Acta Biol. Venezuelica, 18: 7, noted that the only specimen from Venezuela is a juvenile from Temiche, and suggested identifying this from other members of Rhinella is sufficiently difficult as to require additional verification. Barrio-Amorós, 2004, Rev. Ecol. Latino Am., 9: 3, noted Venezuelan distribution and relevant literature, as well as confusion in the literature with Rhinella nasica. Fouquet, Gaucher, Blanc, and Vélez-Rodriguez, 2007, Zootaxa, 1663: 18, excluded this species from the Rhinella margaritifera group. Fenolio, Mendelson, and Lamar, 2012, S. Am. J. Herpetol., 7: 9–15, rediagnosed the species and provided additional data on adult variation, ecology, and range. Rojas-Runjaic, Camargo-Siliet, Carvalho, and La Marca, 2017, Check List, 13(1: 2035): 1–6, reported on new records from Venezuela and mapped the known range. Acosta-Galvis and Señaris, 2018, Vol. 6, Fauna Silvestre Escudo Guayanés: 86, noted the species in Guaviare and Caquetá Provinces, Colombia. See Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 15, for remarks on taxonomy, range, and literature. Señaris and Rojas-Runjaic, 2020, in Rull and Carnaval (eds.), Neotrop. Divers. Patterns Process.: 571–632, commented on range and conservation status in the Venezuelan Guayana. Pereira, Carvalho, Almeida, Rojas-Zamora, Gordo, Frazão, Ávila, and Menin, 2021, Herpetol. Notes, 14: 435–438, mapped records from western Amazonas, Brazil, and discussed the range. Gagliardi-Urrutia, García Dávila, Jaramillo-Martinez, Rojas-Padilla, Rios-Alva, Aguilar-Manihuari, Pérez-Peña, Castroviejo-Fisher, Simões, Estivals, Guillen Huaman, Castro Ruiz, Angulo Chávez, Mariac, Duponchelle, and Renno, 2022, Anf. Loreto: 44–45, provided a brief account, dot map, and genetic barcode for Loreto, Peru.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.