Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhaebo nasicus (Werner, 1903)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhaebo > Species: Rhaebo nasicus

Bufo nasicus Werner, 1903, Zool. Anz., 26: 252. Holotype: IRSNB I.G. 9422 reg. 1.015 (formerly 4792) according to Smith and Laurent, 1950, Bull. Mus. R. Hist. Nat. Belg., 26: 1; subsequently reported as IRSNB 1.015 by Lang, 1990, Doc. Trav., Inst. R. Sci. Nat. Belg., 59: 7. Type locality: "ohne irgend welche Fundortsangabe" (= without locality data); restricted to "South America, probably along the Atlantic drainage" by Smith and Laurent, 1950, Bull. Mus. R. Hist. Nat. Belg., 26: 2.

Rhinella nasica — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 366.

Rhaebo nasicus — Frost, 2006, Amph. Spec. World, vers. 4.0: 434.

English Names

Werner's Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 42).

Distribution

Eastern Venezuela and northwestern Guyana, 100–1410 m elevation.

Geographic Occurrence

Natural Resident: Guyana, Venezuela

Comment

Hoogmoed, 1977, Zool. Meded., Leiden, 51: 265-275, provided an account (as Bufo nasicus). Barrio-Amorós, 1999 "1998", Acta Biol. Venezuelica, 18: 10, commented on the range in Venezuela. Transferred to the Bufo guttatus group (now Rhaebo) by Pramuk, 2006, Zool. J. Linn. Soc., 146: 434, which is Rhaebo of this catalogue. Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 128–129, provided an account. MacCulloch and Lathrop, 2009, R. Ontario Mus. Contrib. Sci., 4: 12, commented on specimens from Mount Ayanganna, Guyana. See Cole, Townsend, Reynolds, MacCulloch, and Lathrop, 2013, Proc. Biol. Soc. Washington, 125: 378–379, for brief account and records for Guyana. Señaris, Lampo, Rojas-Runjaic, and Barrio-Amorós, 2014, Guía Ilust. Anf. Parque Nac. Canaima: 64–65, provided a brief account for the Parque Nacional de Canaima, Venezuela, and photograph. See Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 14–15, for remarks on taxonomy, range (including a record from Auyant-tepui), and literature. Señaris and Rojas-Runjaic, 2020, in Rull and Carnaval (eds.), Neotrop. Divers. Patterns Process.: 571–632, commented on range and conservation status in the Venezuelan Guayana.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.