Rhinella humboldti (Gallardo, 1965)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella humboldti

Bufo granulosus humboldti Gallardo, 1965, Bull. Mus. Comp. Zool., 134: 117. Holotype: MCZ 24882, by original designation. Type locality: "Gualanday, W. of Girardot, Dept. Tolima, Colombia".

Bufo humboltiRivas-Fuenmayor and Barrio-Amorós, 2005, Herpetol. Rev., 36: 205. Incorrect subsequent spelling and undiscussed taxonomic change.

Rhinella humboldtiPramuk, Robertson, Sites, and Noonan, 2008, Global Ecol. Biogeograph., 17: 76; Narvaes and Rodrigues, 2009, Arq. Zool., São Paulo, 40: 43.

English Names

Rivero's Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 40).

Distribution

Colombia in the Magdalena Valley north to the Caribbean and on the eastern side of the Andes in Casanare, Colombia. 

Comment

Narvaes and Rodrigues, 2009, Arq. Zool., São Paulo, 40: 1–73, revised the Rhinella granulosa group, provided an account for this species, and considered Rhinella beebei to be a synonym. Murphy, Angarita-Sierra, Downie, and Jowers, 2017, Salamandra, 53: 267–278, redelimited the species as comprised of populations east of the Andes formerly assigned to Rhinella humboldtiAvila-Pires, Hoogmoed, and Rocha, 2010, Bol. Mus. Parense Emilio Goeldi, Cienc. Nat., 5: 24, suggested that this taxon might not be distinct from some undisclosed member of the Rhinella granulosa group but did not provide any evidence beyond their difficulty in identifying animals. Acosta-Galvis, 2012, Biota Colomb., 13: 213–231, reported the species in the Departamento Sucre, northern Colombia, and briefly discussed habitat. Torres-Suárez and Vargas-Salinas, 2013, Zootaxa, 3702: 198–200, reported on the advertisement call. See account, photograph, and dot map for Colombia by Torres-Suárez and Vargas-Salinas, 2014, Catal. Anf. Rept. Colombia, Medellín, 2(2): 19–23, in the sense of including Rhinella beebeiSimon, Machado, and Marroig, 2016, Proc. R. Soc. London, Ser. B, Biol. Sci., 283 (20161783): 1–10, reported on skull evolution within the Rhinella granulosa group and provided a dot map. Acosta-Galvis, 2017, Biota Colomb., 18: 282–315, reported the species from the Municipality of Yopal, Casanare Department, Colombia. In the Rhinella marina clade, Rhinella granulosa group of Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1–156. Rivera, Prates, Caldwell, Rodrigues, and Fujita, 2022, Heredity, 130: 14–21, reported on mtDNA phylogenetics in the Rhinella granulosa group, including this species.   

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