Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhinella humboldti (Gallardo, 1965)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella humboldti

Bufo granulosus humboldti Gallardo, 1965, Bull. Mus. Comp. Zool., 134: 117. Holotype: MCZ 24882, by original designation. Type locality: "Gualanday, W. of Girardot, Dept. Tolima, Colombia".

Bufo humbolti — Rivas-Fuenmayor and Barrio-Amorós, 2005, Herpetol. Rev., 36: 205. Incorrect subsequent spelling and undiscussed taxonomic change.

Rhinella humboldti — Pramuk, Robertson, Sites, and Noonan, 2008, Global Ecol. Biogeograph., 17: 76; Narvaes and Rodrigues, 2009, Arq. Zool., São Paulo, 40: 43.

English Names

Rivero's Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 40).

Distribution

Colombia in the Magdalena Valley north to the Caribbean and on the eastern side of the Andes in Casanare, Colombia.

Comment

Narvaes and Rodrigues, 2009, Arq. Zool., São Paulo, 40: 1–73, revised the Rhinella granulosa group, provided an account for this species, and considered Rhinella beebei to be a synonym. Murphy, Angarita-Sierra, Downie, and Jowers, 2017, Salamandra, 53: 267–278, redelimited the species as comprised of populations east of the Andes formerly assigned to Rhinella humboldti. Avila-Pires, Hoogmoed, and Rocha, 2010, Bol. Mus. Parense Emilio Goeldi, Cienc. Nat., 5: 24, suggested that this taxon might not be distinct from some undisclosed member of the Rhinella granulosa group but did not provide any evidence beyond their difficulty in identifying animals. Acosta-Galvis, 2012, Biota Colomb., 13: 213–231, reported the species in the Departamento Sucre, northern Colombia, and briefly discussed habitat. Torres-Suárez and Vargas-Salinas, 2013, Zootaxa, 3702: 198–200, reported on the advertisement call. See account, photograph, and dot map for Colombia by Torres-Suárez and Vargas-Salinas, 2014, Catal. Anf. Rept. Colombia, Medellín, 2(2): 19–23, in the sense of including Rhinella beebei. Simon, Machado, and Marroig, 2016, Proc. R. Soc. London, Ser. B, Biol. Sci., 283 (20161783): 1–10, reported on skull evolution within the Rhinella granulosa group and provided a dot map. Acosta-Galvis, 2017, Biota Colomb., 18: 282–315, reported the species from the Municipality of Yopal, Casanare Department, Colombia. In the Rhinella marina clade, Rhinella granulosa group of Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1–156. Rivera, Prates, Caldwell, Rodrigues, and Fujita, 2022, Heredity, 130: 14–21, reported on mtDNA phylogenetics in the Rhinella granulosa group, including this species.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.