Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhinella leptoscelis (Boulenger, 1912)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella leptoscelis

Bufo leptoscelis Boulenger, 1912, Ann. Mag. Nat. Hist., Ser. 8, 10: 186. Holotype: BMNH 1907.5.7.32 (reregistered 1947.2.21.95), according to Hoogmoed, 1985, in Frost (ed.), Amph. Species World: 51, and museum records. Type locality: "Santo Domingo, Carabaya, S.E. Peru, 6500 feet".

Rhinella leptoscelis — Padial, Chaparro Auza, Köhler, and De la Riva, 2009, Zootaxa, 2115: 56.

Common Names

None noted.

Distribution

Humid forests of the Amazonian versant of the Andes from central Bolivia (Cochabamba and La Paz) to southern Peru (Cusco and Puno), 1300–1658 m elevation.

Geographic Occurrence

Natural Resident: Bolivia, Peru

Comment

Removed from the synonymy of Rhinella veraguensis by Padial, Chaparro Auza, Köhler, and De la Riva, 2009, Zootaxa, 2115: 56–64, where it had been placed by Hoogmoed, 1990, in Peters and Hutterer (eds.), Vert. Tropics: 116; although this synonymy was disputed implicitly by Duellman and Schulte, 1992, Copeia, 1992: 162–172, but who provided no additional evidence. In the Rhinella margaritifera clade, Rhinella veraguensis group of Pereyra, Blotto, Baldo, Chaparro Auza, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1–156. Köhler, Vences, Padial, Plewnia, and Lötters, 2023, Salamandra, 59: 321, provided a dot map of the species and noted the change in range of this species with the exclusion of Rhinella kuka. Ocampo, Aparicio, Bernal Hoverud, Domic, and Wallace, 2024, Herpetol. Notes, 17: 371–389, reported the species and its habitat in Madidi National Park, La Paz, Bolivia. Vargas, Jaico, and Catenazzi, 2026, Zootaxa, 5884: 97–118, reported the species from Huancavelica Department, Peru, at 2165 m elevation.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist