Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhinella quechua (Gallardo, 1961)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella quechua

Bufo quechua Gallardo, 1961, Breviora, 141: 4. Holotype: CM 4225, by original designation. Type locality: "Incachaca, 2500 m, Department of Cochabamba, Bolivia".

Bufo echinodes Reynolds and Foster, 1992, Herpetol. Monogr., 6: 87. Holotype: USNM 257799, by original designation. Type locality: "Parjacti [= Paracti], 83.2 km by road NE Cochabamba on road to Villa Tunari, Chapare Province, Department of Cochabamba, Bolivia, elevation approximately 2044 m". Synonymy by Harvey and Smith, 1993, Proc. Biol. Soc. Washington, 106: 442–449. "Incachaca, 2500 m, Department of Cochabamba, Bolivia".

Bufo echinoides — Harvey and Smith, 1993, Proc. Biol. Soc. Washington, 106: 442–449. Incorrect subsequent spelling.

Chaunus quechua — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 364.

Rhinella quechua — Chaparro, Pramuk, and Gluesenkamp, 2007, Herpetologica, 63: 205.

English Names

Incachaca Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 43).

Distribution

Eastern slopes of the Bolivian Andes from Departaments of Cochabamba, Verdecillo, and Santa Cruz, 1900–2600 m elevation.

Geographic Occurrence

Natural Resident: Bolivia

Endemic: Bolivia

Comment

In the Bufo ockendeni (= Rhinella veraguensis) group in the original publication, a group recognized by Cei, 1980, Monit. Zool. Ital., N.S., Monogr., 2: 203. Although asserted to be a member of Rhinella (as Bufo typhonius group) group by Blair (ed.), 1972, Evol. Genus Bufo, its membership in the Rhinella veraguensis group (as Bufo) was supported by Hoogmoed, 1990, in Peters and Hutterer (eds.), Vert. Tropics: 117, and Harvey and Smith, 1993, Proc. Biol. Soc. Washington, 106: 442–449. See Lavilla and Cei, 2001, Monogr. Mus. Reg. Sci. Nat. Torino, 28: 26–27, noted confusion in the literature with Rhinella rumbolli. See brief accounts (as both Bufo amboroensis and Bufo quechua) by Köhler, 2000, Bonn. Zool. Monogr., 48: 82, 86–87. See photograph, map, description of geographic range and habitat, and conservation status (as Bufo quechua) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 190. Sosa, Schalk, Braga, and Pinto-Ledezma, 2015, Herpetol. Rev., 46: 214, provided a record (as Rhinella amboroensis) from Verdecillo, Province of Manuel Maria Caballero, Santa Cruz, Bolivia. In the Rhinella margaritifera clade, Rhinella veraguensis group of Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1–156. See comment under Rhinella amboroensis. Köhler, Vences, Padial, Plewnia, and Lötters, 2023, Salamandra, 59: 321, provided a dot map of the species.

External links:

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For additional sources of general information from other websites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.