Rhinella vellardi (Leviton and Duellman, 1978)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella vellardi

Bufo spinulosus orientalis Vellard, 1959, Mem. Mus. Hist. Nat. Javier Prado, 8: 1-48. Syntypes: MUSM (formerly MHNJP) 815 (2 specimens); MUSM (formerly MHNJP) 3750 (renumbered) designated lectotype by Morales, Carrillo, and Ortega, 1990, Publ. Mus. Hist. Nat. Univ. Nac. Mayor San Marcos, 33: 2. Type locality: "Hacienda Yanasona (Alto Marañón...)", Peru. Preoccupied by Bufo viridis orientalis Werner, 1896 "1895".

Bufo limensis orientalisCei, 1972, in Blair (ed.), Evol. Genus Bufo: 82-91.

Bufo orientalisCei, 1972, in Blair (ed.), Evol. Genus Bufo: 82-91. Alternative arrangement.

Bufo vellardi Leviton and Duellman, 1978, J. Herpetol., 12: 246. Replacement name for Bufo spinulosus orientalis Vellard, 1959.

Bufo limensis vellardiLeviton and Duellman, 1978, J. Herpetol., 12: 246. Alternative arrangement.

Chaunus vellardiFrost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 364. Savage and Bolaños, 2009, Zootaxa, 2005: 4, by implication.

Rhinella vellardiChaparro, Pramuk, and Gluesenkamp, 2007, Herpetologica, 63: 211, by implication; Pramuk, Robertson, Sites, and Noonan, 2008, Global Ecol. Biogeograph., 17: 76.

English Names

Alto Maranon Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 44).


Known only from the type locality in the upper Marañón area, Cajamarca, northern Peru.

Geographic Occurrence

Natural Resident: Peru

Endemic: Peru


Considered a subspecies of Rhinella limensis (as Bufo limensis) by Leviton and Duellman, 1978; see comment under Rhinella limensisVera Candioti, Grosso, Pereyra, Haad, Lescano, Siu-Ting, Aguilar, and Baldo, 2020, Herpetol. Monogr., 34: 116–130, reported on larval morphology who considered it to be in the Rhinella spinulosa group. in the Rhinella marina clade, Rhinella spinulosa group of Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1– 155. 

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