Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhinella veraguensis (Schmidt, 1857)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Rhinella > Species: Rhinella veraguensis

Bufo veraguensis Schmidt, 1857, Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 24: 10. Holotype: KM 1032/1350, lost, according to Savage, 1970, Proc. California Acad. Sci., Ser. 4, 38: 273–288. BMNH 1947.2.21.23 (lectotype of Bufo ockendeni), designated neotype by Hoogmoed, 1990, in Peters and Hutterer (eds.), Vert. Tropics: 116. Type locality: "Neu-Granada"; subsequently stated by Schmidt, 1858, Denkschr. Akad. Wiss. Wien, Math. Naturwiss. Kl., 14: 254, as "Neu-Granada, Provinz Veragua" in error according to Savage, 1969, Copeia, 1969: 178–179, who suggested that the type came from Bolivia or Peru. Neotype from "Marcapata Valley, E. Peru". See discussion by Savage, 1970, Proc. California Acad. Sci., Ser. 4, 38: 273–288.

Chilophryne veraguensis — Cope, 1862, Proc. Acad. Nat. Sci. Philadelphia, 14: 358.

Incilius veraguensis — Cope, 1863, Proc. Acad. Nat. Sci. Philadelphia, 15: 50.

Bufo ockendeni Boulenger, 1902, Ann. Mag. Nat. Hist., Ser. 7, 10: 396. Syntypes: BMNH (several specimens), by original designation, these being BMNH 1947.2.21.2637 (formerly 1902.5.29.130-131) and 1947.2.21.21–25 (formerly 1902.5.29.199–203), according to museum records. BMNH 1947.2.21.23 designated lectotype by Savage, 1969, Copeia, 1969: 179. Type locality: "Marcapata Valley, E. Peru" and "Charuplaya, Bolivia"; restricted to "Marcapata Valley, E. Peru" by lectotype designation. Synonymy by Savage, 1969, Copeia, 1969: 178–179.

Chaunus veraguensis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 364.

Rhinella veraguensis — Chaparro, Pramuk, and Gluesenkamp, 2007, Herpetologica, 63: 205.

Common Names

Veragua Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 44).

Distribution

Southeastern Peru and the departments of Cochabamba, Chuquisaca, La Paz, and Santa Cruz, Bolivia, between 1300 and 1900 m, in interandean valleys and the Amazonian versant.

Geographic Occurrence

Natural Resident: Bolivia, Peru

Comment

Removed from Rhinella (as the Bufo margaritifer group) by Hoogmoed, 1990, in Peters and Hutterer (eds.), Vert. Tropics: 116. See comment under Rhinella inca. In the Rhinella veraguensis group (as Bufo) of Duellman and Schulte, 1992, Copeia, 1992: 162–172. Cadle and Altig, 2008, Stud. Neotrop. Fauna Environ., 26: 45–53, described larval morphology. See brief account by Köhler, 2000, Bonn. Zool. Monogr., 48: 87. De la Riva and Reichle, 2014, Herpetol. Monogr., 28: 59, who discussed conservation status in Bolivia, noted that this species has become very rare in that country. In the Rhinella margaritifera clade, Rhinella veraguensis group of Pereyra, Blotto, Baldo, Chaparro, Ron, Elias-Costa, Iglesias, Venegas, Thomé, Ospina-Sarria, Maciel, Rada, Kolenc, Borteiro, Rivera-Correa, Rojas-Runjaic, Moravec, De la Riva, Wheeler, Castroviejo-Fisher, Grant, Haddad, and Faivovich, 2021, Bull. Am. Mus. Nat. Hist., 447: 1–156. Köhler, Vences, Padial, Plewnia, and Lötters, 2023, Salamandra, 59: 321, provided a polygon map of the species.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.