Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Werneria mertensiana Amiet, 1976

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Werneria > Species: Werneria mertensiana

Bufo mertensi Amiet, 1972, Ann. Fac. Sci. Cameroun, 11: 125. Holotype: JLA 70.380, by original desgnation; now MHNG 1253.91 according to Grandison, 1985, in Frost (ed.), Amph. Species World: 77; and Schätti, Perret, and Mariaux, 2002, Cat. Comm. Types Amph. Rept. Mus. Hist. Nat. Geneve, Vers. 4.0 (Jan. 2002): 20. Type locality: "Mt Nlonako, env. de N'Kongsamba, 1.000 m", Cameroon. Junior homonym of Bufo ictericus mertensi Cochran, 1950.

Werneria mertensiana Amiet, 1976, Rev. Zool. Bot. Afr., 90: 43. Replacement name for Bufo mertensi Amiet, 1972.

Bufo mertensiana — Grandison, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 187-215; Lawson, 1993, Herpetol. Nat. Hist., 1: 41.

Werneria preussi mertensiana — Amiet, 1982, Mem. Mus. Natl. Hist. Nat. Paris, A—Zool., 132: 29-42; De la Riva, 1994, Rev. Esp. Herpetol., 8: 129.

Common Names

Mertens' Smalltongue Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 46).

Mertens' Torrent Toad (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 100).

Distribution

Western Cameroonian mountain areas, to the Kala and Mbam–Minkoum moutnains around Yaoundé, southern Cameroon, as well arguably as from Monte Alén in Equatorial Guinea (see comment), 800 to 1050 m elevation.

Geographic Occurrence

Natural Resident: Cameroon

Likely/Controversially Present: Equatorial Guinea

Endemic: Cameroon

Comment

In a privately circulated "List provisoire des amphibiens anoures du Cameroun" J.-L. Amiet suggested that this is a subspecies of Werneria preussi. Rödel, Schmitz, Pauwels, and Böhme, 2004, Zootaxa, 720: 1-28, discussed confusion with other species. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 206. Rödel, Schmitz, Pauwels, and Böhme, 2004, Zootaxa, 720: 23, discussed the Equatorial Guinea record. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 100–101, provided a brief account, photograph, and range map. See brief account (as Werneria cf. mertensiana), range map, and photographs for Equatorial Guinea in Sánchez-Vialas, Calvo-Revuelta, Castroviejo-Fisher, and De la Riva, 2020, Proc. California Acad. Sci., Ser. 4, 66: 137–230.

External links:

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For access to general information see Wikipedia
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.