Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Werneria preussi (Matschie, 1893)

Class: Amphibia > Order: Anura > Family: Bufonidae > Genus: Werneria > Species: Werneria preussi

Bufo preussi Matschie, 1893, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1893: 175. Syntypes: ZMB (originally 60 specimens), BMNH 1935.2.8.3 (now 1947.2.20.43) and 1898.9.14.7 (now 1947.2.20.42) by museum records (presumably exchanged from ZMB), MCZ 10296 (exchanged from ZMB according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 232), according to Grandison, 1985, in Frost (ed.), Amph. Species World: 77. ZMB 93453 designated lectotype by Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 361. Type locality: "Buea, Kamerun, 950 m".

Atelopus africanus Werner, 1898, Verh. Zool. Bot. Ges. Wien, 48: 201. Syntypes: ZMB, and including NHMW 20995, according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 13, Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 17, and Gemel, Gassner, and Schweiger, 2019, Ann. Naturhist. Mus. Wien, Ser. B, 121: 45. Type locality: "Bismarkburg (Adeli, Togo)", in error according to Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 311, and Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 351. Synonymy by Boulenger, 1899, Zool. Rec., 35: 27; Nieden, 1923, Das Tierreich, 46: 105; Perret, 1966, Zool. Jahrb., Jena, Abt. Syst., 93: 311.

Stenoglossa fulva Andersson, 1903, Verh. Zool. Bot. Ges. Wien, 53: 144. Holotype: NHMG, although not listed in current type list. Type locality: "Buca im Kamerungebirge in einem Regenbach" (= Cameroon range [Buea]). Synonymy by Tandy and Keith, 1972, in Blair (ed.), Evol. Genus Bufo: 155.

Werneria fulva — Poche, 1903, Zool. Anz., 26: 701.

Bufo (Palaeobufo) reussi — Bolkay, 1919, Glasn. Zemaljskog Muz. Bosni Hercegov., 31: 295. Error for "preussi" according to Dubois, 2019, Alytes, 37: 75.

Werneria preussi — Amiet, 1976, Rev. Zool. Bot. Afr., 90: 43; Grandison, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 187–215.

Werneria preussi preussi — Amiet, 1982, Mem. Mus. Natl. Hist. Nat. Paris, A—Zool., 132: 29–42.

Werneria submontana Rödel, Schmitz, Pauwels, and Böhme, 2004, Zootaxa, 720: 17. Holotype: ZFMK 69699, by original designation. Type locality: "Bakossi Mts., Edib Hills, Lake Edib, N 4° 57′, E 9° 39′, 1,100 m a.s.l., Cameroon". Synonymy by Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 351.

Werneria africanus — Segniagbeto, Bowessidjaou, Dubois, and Ohler, 2007, Alytes, 24: 83. Conjectural combination. Gender disagreement.

Werneria preussi preussi — Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 351.

Werneria preussi submontana — Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 351.

Common Names

Buea Smalltongue Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 46).

Preuss' Torrent Toad (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 98).

Hill Torrent Toad (Werneria submontana: Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 100).

Distribution

Bakossi Mts., Mwendelengo Mts., and Nyasoso at the base of Mt. Kupe, Cameroon Highlands, 800-1200 m elevation, occasionally as low as 200 m; n creeks of the highlands (700–1200 m elevation) of western Cameroon on the lower slopes of Mount Cameroon and Mount Kupe, West Africa.

Geographic Occurrence

Natural Resident: Cameroon

Likely/Controversially Present: Cameroon

Comment

Grandison, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 187–215, discussed characters and relationships of this species. See also accounts by Mertens, 1938, Abh. Senckenb. Naturforsch. Ges., 442: 10, and Amiet, 1972, Ann. Fac. Sci. Cameroun, 11: 121–140. Sole member of the Bufo preussi group of Tandy and Keith, 1972, in Blair (ed.), Evol. Genus Bufo: 155. Record for Equatorial Guinea by De la Riva, 1994, Rev. Esp. Herpetol., 8: 123–138. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 162–163, reported on comparative tadpole morphology. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 98–99 (as Werneria preussi) and 100–101 (as Werneria submontana), provided brief accounts, photographs, and range maps. See (as Werneria submontana) photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 206. Petzold, Norden, Gonwouo, Segniagbeto, Ohler, and Rödel, 2025, Zootaxa, 5575: 351–373, discussed the phylogenetics and systematics of the species.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist