Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

"Colostethus" ruthveni Kaplan, 1997

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Dendrobatidae > Subfamily: Dendrobatinae > Species: "Colostethus" ruthveni

Colostethus ruthveni Kaplan, 1997, J. Herpetol., 31: 369. Holotype: UMMZ 54618, by original designation. Type locality: "Quebrada Viernes Santo, in the mountains near Santa Marta, Departamento de Magdalena, Colombia".

"Colostethus" ruthveni — Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado, and Rueda-Almonacid, 2017, S. Am. J. Herpetol., 12 (Special Issue): 26.

English Names

None noted.

Distribution

Elevations of 472–2100 m on the Sierra Nevada de Santa Marta, Departamento del Magdalena, Colombia. See comment.

Geographic Occurrence

Natural Resident: Colombia

Endemic: Colombia

Comment

See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 226. González-Maya, González, Zárrate-Charry, Charry, Cepeda, and Balaguera-Reina, 2011, J. Threatened Taxa, 3: 1633–1636, provided a range extension, as did Granda-Rodríguez, Saboyá-Acosta, del Portillo-Mozo, and Renjifo, 2014, Check List, 10: 674–676, wo provided a range extension and mapped the known localities of this species as well as providing photographs. Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado, and Rueda-Almonacid, 2017, S. Am. J. Herpetol., 12 (Special Issue): 1–90, noted that the Colostethus ruthveni group (Colostethus ruthveni and an unnamed species) sits within Dendrobatinae and not within Colostethus. Pending a generic description by Grant et al. I retain "Colostethus" ruthveni in Dendrobatinae. Meza-Joya, Chinchilla-Lemus, Ramos, Armesto, and Acevedo-Rincón, 2019, Herpetol. Notes, 12: 61–69, provided new localities, provided a dot map, and discussed the range, identification, and taxonomic affinities. Jiménez-Bolaño, Montes-Correa, Polo-Córdoba, Linares-Vargas, Vergara-Gil, Barrio-Amorós, and Koch, 2019, Salamandra, 55: 27–38, reported on the acoustic repertoire of the group and corroborated the conclusion of Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado, and Rueda-Almonacid, 2017, S. Am. J. Herpetol., 12 (Special Issue): 1–90, that the population from the southern slopes of the Sierra Nevada de Santa Marta are a distinct, still unnamed, species (referred to by Grant et al. as “Colostethus” sp. ruthveni-like). They further suggested that “Colostethus” ruthveni sensu stricto is restricted to the northwestern slopes Sierra Nevada de Santa Marta, with specimens from the northern slopes representing an additional, undescribed species, referred to by them as “Colostethus” cf. ruthveni. The claim by Portik, Streicher, and Wiens, 2023, Mol. Phylogenet. Evol., 188 (107907): 67, that their placement of Colostethus ruthveni in Dendrobatinae renders Colostethinae non-monophyletic is surprising inasmuch as Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado, and Rueda-Almonacid, 2017, S. Am. J. Herpetol., 12 (Special Issue): 1–90 (not cited by Portik et al., 2023) already noted that the Colostethus ruthveni group (Colostethus ruthveni and an unnamed species) sits within Dendrobatinae and not within Colostethus. Grant et al. (2017) placed quotation marks around Colostethus to mark that its generic assignment remains to be identified.

External links:

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For additional sources of general information from other websites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.