Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyla carthaginiensis Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, and Litvinchuk, 2019

Class: Amphibia > Order: Anura > Family: Hylidae > Genus: Hyla > Species: Hyla carthaginiensis

Hyla carthaginiensis Dufresnes, Beddek, Skorinov, Fumagalli, Perrin, Crochet, and Litvinchuk, 2019, Mol. Phylogenet. Evol., 134: 296. Holotype: ZISP 13722, by original designation. Type locality: "near Qusur Lireba, 10 km SSE of Tabarka, Tunisia (36.868°N, 8.728°E, 113m a.s.l.)". Zoobank publication registration: 16C1D704-8906-4E84-A0D1-345FF9F8A664

Hyla numidica Escoriza and Ben Hassine, 2019, Amph. N. Afr.: 190. Types: not explicitly designated. Type locality: "Nefza (Tunisia)”. Unavailable name as discussed by Dufresnes and Crochet, 2020, Zootaxa, 4759: 597–599.

Common Names

Carthaginian Tree Frog (original publication).

Distribution

Humid Mediterranean regions of northeastern Algeria and northern Tunisia.

Geographic Occurrence

Natural Resident: Algeria, Tunisia

Comment

The sister taxon of Hyla meridionalis according to the original publication. Stöck, Dufresnes, Litvinchuk, Lymberakis, Biollay, Berroneau, Borzée, Ghali, Ogielska, and Perrin, 2012, Mol. Phylogenet. Evol., 65: 1–9, reported on the genetic diversity among populations and provided a dot map of the range (including their "Hyla new sp. 1", subsequently named as Hyla carthaginiensis. See Dufresnes and Crochet, 2020, Zootaxa, 4759: 597–599, for range map. Dufresnes, Berroneau, Dubey, Litvinchuk, and Perrin, 2020, Sci. Rep. (Nature, London), 10 (5502): 1–10, discussed the genetics of the contact zone of this species with Hyla meridionalis. Nouira, Blanc, Crochet, Frétey, Géniez, Ineich, De Massary, Ohler, Tlili, and Lescure, 2022, Bull. Soc. Herpetol. France, 180: 12, discussed the species in Tunisia.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist