Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dendropsophus columbianus (Boettger, 1892)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dendropsophus > Species: Dendropsophus columbianus

Hyla columbiana Boettger, 1892, Kat. Batr. Samml. Mus. Senckenb. Naturforsch. Ges.: 41. Syntypes: SMF 1375.2s (2 specimens); SMF 2365 designated lectotype by Mertens, 1967, Senckenb. Biol., 48(A): 41. Type locality: "Popayán, Ver. Staaten von Columbia", Departamento Cauca, Colombia.

Hyla variabilis Boulenger, 1896, Ann. Mag. Nat. Hist., Ser. 6, 17: 20. Syntypes: BMNH 1947.2.13.15–31 (formerly 1895.11.16.65–81 according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 96), NHMW 19438.1–3 (according to Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 20, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 24), MZUT An337 (3 specimens) (according to Gavetti and Andreone, 1993, Cat. Mus. Reg. Sci. Nat., Torino, 10: 77), FMNH 3565, MCZ 2606 (according to Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 280), UMMZ 46464, 51269, 58908 (8 specimens; according to Peters, 1952, Occas. Pap. Mus. Zool. Univ. Michigan, 539: 17), USNM 71115 (according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 60); ZMH A00963 according to Hallermann, 1998, Mitt. Hamburg. Zool. Mus. Inst., 95: 199; ZSM 1182/0/1–3 according to Glaw and Franzen, 2006, Spixiana, München, 29: 170; ZFMK 28635, according to Böhme, 2014, Mertensiella, 21: 101, and RMNH 4496, according to Gassó Miracle, van den Hoek Ostende, and Arntzen, 2007, Zootaxa, 1482: 45. Type locality: "Cali", Colombia. Synonymy by Duellman and Trueb, 1983, in Rhodin and Miyata (eds.), Adv. Herpetol. Evol. Biol.: 33–51.

Dendropsophus columbianus — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 90.

English Names

Boettger's Colombian Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 55).

Distribution

Western slope of the Cordillera Central and easter slope of the Cordillera Occidental (Departments of Nariño, Cauca, Quindío, Risaralda, and Valle del Cauca), Colombia, 950–2300 m elevation.

Comment

In the Dendropsophus columbianus group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 90. See account, photograph, and dot map by Agudelo-Valderrama, Bolívar-García, and Hernández-Medina, 2014, Catal. Anf. Rept. Colombia, Medellín, 2(1): 40–46. Ramírez-Chaves, Pisso-Florez, Liévano-Bonilla, Ayerbe-Quiñones, Anganoy-Criollo, and Noguera-Urbano, 2018, Herpetol. Notes, 11: 49–58, discussed the range of the species in Colombia with new records and provided a dot map as well as a predicted distribution map. In the Dendropsophus columbianus group of Orrico, Grant, Faivovich, Rivera-Correa, Rada, Lyra, Cassini, Valdujo, Schargel, Machado, Wheeler, Barrio-Amorós, Loebmann, Moravec, Zina, Solé, Sturaro, Peloso, Suárez, and Haddad, 2021, Cladistics, 37: 73–105, whose samples of Dendropsophus columbianus were paraphyletic with respect to Dendropsophus bogerti, something the authors suggested required additional investigation.

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