Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dendropsophus molitor (Schmidt, 1857)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dendropsophus > Species: Dendropsophus molitor

Hyla molitor Schmidt, 1857, Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 24: 11. Syntypes: Including NHMW 16494 (according to Duellman, 1977, Das Tierreich, 95: 77, and Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 19), KM 1010/1341 (2 specimens); female of KM syntypes designated lectotype by Savage and Heyer, 1969, Rev. Biol. Tropical, 16: 1–127. Type locality: "Chiriquiflusse unweit Bocca del toro", Panama; in error, according to Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 650.

Hyla molitor var. marmorata Schmidt, 1857, Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 24: 12. Holotype: KM 1010/1342, according to Savage, 1970, Proc. California Acad. Sci., Ser. 4, 38: 273–288. Type locality: "Chiriqui-Flusse unweit Bocca del toro", Panama. Preoccupied by Hyla marmorata Laurenti, 1768. Synonymy by Jungfer, 2017, Salamandra, 53: 20.

Hyla labialis Peters, 1863, Monatsber. Preuss. Akad. Wiss. Berlin, 1863: 463. Holotype: ZMB 4913, according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 42. Type locality: "Umgegend von Bogotá" (= vicinity of Bogota), Departamento Cundinamarca, Colombia.: 77. Synonymy by Jungfer, 2017, Salamandra, 53: 20.

Hyla creolica Werner, 1899, Verh. Zool. Bot. Ges. Wien, 49: 483. Syntypes: BMNH 1947.2.24.14 (formerly 1900.2.7.5; according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 89) and (formerly ZIUG) ZFMK 28569–74 (Chapinero near Bogota), 28575 (Savanna near Bogota), 28576–77 (Boqueron near Bogota), 28578–79 (Porero near Bogota), 28580–83 (Hacienda Otoño near Bogota), according to Böhme and Bischoff, 1984, Bonn. Zool. Monogr., 19: 181, and Böhme, 2014, Mertensiella, 21: 100. Type localities: "Savanna bei Bogotá" and "Hacienda Otoño bei Bogotá", Department of Cundinamarca, Colombia. Synonymy with Hyla vilsoniana by Boulenger, 1900, Zool. Rec., 36: 29 (who documented that Hyla vilsoniana antedated Hyla creolica by seven months); Peracca, 1914, Mem. Soc. Neuchatel. Sci. Nat., 5: 108, and Hellmich, 1940, Zool. Anz., 129: 1. Synonymy by Dunn, 1944, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 6: 77. Not mentioned in synonymy of Jungfer, 2017, Salamandra, 53: 20.

Hyla servalina Werner, 1899, Verh. Zool. Bot. Ges. Wien, 49: 483. Holotype: ZFMK 28584 (formerly ZIUG) according to Böhme and Bischoff, 1984, Bonn. Zool. Monogr., 19: 182. Type locality: "Boquéron bei Bogotá", Department of Cundinamarca, Colombia. Synonymy by Dunn, 1944, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 6: 77. Not mentioned in synonymy of Jungfer, 2017, Salamandra, 53: 20.

Hyla vilsoniana Cope, 1899, Philadelphia Mus. Sci. Bull., 1: 3. Syntypes: "eight specimens, Nos. 75, 76, and 87", althgouh measurements given for only one specimen, implying a holotype and paratypes were intended; according to museum records AMNH 5456–61, 53822–24 according to Duellman, 1977, Das Tierreich, 95: 66. Type locality: "neighborhood of Bogota", Department of Cundinamara, Colombia. Synonymy by Dunn, 1944, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 6: 68–81. Not mentioned in synonymy of Jungfer, 2017, Salamandra, 53: 20.

Hyla wilsoniana — Peracca, 1914, Mem. Soc. Neuchatel. Sci. Nat., 5: 108. Incorrect subsequent spelling.

Hyla gularis Werner, 1916, Zool. Anz., 47: 302. Syntypes: Brünn Landsmuseum, by original designation; status of specimens currently known. Type locality: "Bogota", Cundinamarca, Colombia. Synonymy by Dunn, 1944, Rev. Acad. Colomb. Cienc. Exact. Fis. Nat., 6: 75. Not mentioned in synonymy of Jungfer, 2017, Salamandra, 53: 20.

Hyla vilsoniana krausi Hellmich, 1940, Zool. Anz., 129: 8. Holotype: ZSM 102/1937, according to Duellman, 1977, Das Tierreich, 95: 66, and Glaw and Franzen, 2006, Spixiana, München, 29: 170. Type locality: "Laguna de la Guitarra (Paramo de Sumapaz)", Department of Cundinamarca, Colombia. Subspecies distinctiveness rejected by Duellman, 1989, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 131: 1–12. Not mentioned in synonymy of Jungfer, 2017, Salamandra, 53: 20.

Hyla labialis labialis — Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 254.

Hyla labialis krausi — Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 254.

Dendropsophus labialis — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 91.

Dendropsophus luddeckei Guarnizo, Escallón, Cannatella, and Amézquita, 2012, Herpetologica, 68: 535. Holotype: ANDES-A 0603, by original designation. Type locality: "Sotaquira, (05°45′02.8″N; 73°16′39.7″W), 2765 m elevation, Departamento de Boyacá, Colombia". Synonymy by Arias-Cárdenas, Barrientos, Pardo-Diaz, Paz, Crawford, and Salazar, 2023, Zool. J. Linn. Soc., https://doi.org/10.1093/zoolinnean/zlad085: 1.

Dendropsophus molitor — Jungfer, 2017, Salamandra, 53: 18.

Common Names

Green Dotted Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 56).

Schmidt's Bolivian Treefrog (Hyla molitor var. marmorata [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 56).

Distribution

Cordillera Oriental of Colombia in the Departments of Cundinamarca, Boyacá, and Norte de Santander, Colombia. See comment.

Geographic Occurrence

Natural Resident: Colombia

Endemic: Colombia

Comment

See Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 251, for account. Removed from the synonymy of Smilisca phaeota by Cochran and Goin, 1970, Bull. U.S. Natl. Mus., 288: 254, where it had been placed by Duellman, 1968, Cat. Am. Amph. Rept., 61: 1–2. Lynch and Renjifo, 2001, Guia Anf. Rept. Bogota: 48–49, presented a brief account and photograph. In the Dendropsophus labialis group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 91. Guarnizo, Amézquita, and Bermingham, 2009, Mol. Phylogenet. Evol., 50: 84–92, reported on molecular phylogeography within eastern Colombia, in what subsequently was named Dendropsophus luddeckei. Guarnizo, Escallón, Cannatella, and Amézquita, 2012, Herpetologica, 68: 523–540, revised and redelimited the species. Guarnizo and Cannatella, 2014, J. Heredity, 20: 216–225, reported on gene flow characteristics. Estupiñán, Ferrari, Gonçalves, Barbosa, Vallinoto, and Schneider, 2016, ZooKeys, 637: 89–106, suggested on the basis of COI barcodes that cryptic species are likely. Jungfer, 2017, Salamandra, 53: 18–24, discussed Hyla molitor Schmidt 1857, and concluded that it is a senior synonym of Hyla labialis Peters, 1863; in his synonymy of Dendropsophus molitor he did not include a number of names traditionally in the synonymy of Denropsophus labialis, although what this implies is unknown. \ In the Dendropsophus molitor group of Orrico, Grant, Faivovich, Rivera-Correa, Rada, Lyra, Cassini, Valdujo, Schargel, Machado, Wheeler, Barrio-Amorós, Loebmann, Moravec, Zina, Solé, Sturaro, Peloso, Suárez, and Haddad, 2021, Cladistics, 37: 73–105. Arias-Cárdenas, Barrientos, Pardo-Diaz, Paz, Crawford, and Salazar, 2023, Zool. J. Linn. Soc., https://doi.org/10.1093/zoolinnean/zlad085: 1–13, discussed molecular markers, morphometrics, advertisement call, comparative morphology and placed Dendropsophus ludduckei into synonymy.

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