Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dendropsophus phlebodes (Stejneger, 1906)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dendropsophus > Species: Dendropsophus phlebodes

Hyla phlebodes Stejneger, 1906, Proc. U.S. Natl. Mus., 30: 817. Holotype: USNM 29970, by original designation. Type locality: "San Carlos, [Boca de Arenal, Cantón de San Carlos, Provincia Alajuela,] Costa Rica". Type locality discussed by Savage, 1974, Rev. Biol. Tropical, 22: 77, who corrected it to "Boca de Arenal, Cantón de San Carlos, Provincia de Alajuela; 55 m", Costa Rica.

Dendropsophus phlebodes — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 92.

Common Names

San Carlos Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 57).

San Carlos Dwarf Treefrog (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 13).

Distribution

Caribbean lowlands from southern Nicaragua to Panama and Pacific lowlands of eastern Panama and northwestern Colombia, 20 to 700 m elevation.

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Nicaragua, Panama

Comment

See Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 220–223, and Savage and Heyer, 1969, Rev. Biol. Tropical, 16: 1–127. Lips and Savage, 1996, Proc. Biol. Soc. Washington, 109: 17–26, included this species (as Hyla phlobodes) in a key to the tadpoles found in Costa Rica. See account by Savage, 2002, Amph. Rept. Costa Rica: 318–320, who placed this species in his Hyla leucophyllata group, Hyla microcephala subgroup. In the Dendropsophus microcephalus group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 91–92. See comments by Sunyer, Páiz, Dehling, and Köhler, 2009, Herpetol. Notes, 2: 189–202, regarding Nicaraguan populations. Köhler, 2011, Amph. Cent. Am.: 216–218, provided a brief summary of natural history of the Central American species, including this one, and provided a range map and photograph. In the Dendropsophus microcephalus group of Orrico, Grant, Faivovich, Rivera-Correa, Rada, Lyra, Cassini, Valdujo, Schargel, Machado, Wheeler, Barrio-Amorós, Loebmann, Moravec, Zina, Solé, Sturaro, Peloso, Suárez, and Haddad, 2021, Cladistics, 37: 73–105. Not addressed in the study of Whitcher, Orrico, Ron, Lyra, Cassini, Ferreira, Nakamura, Peloso, Rada, Rivera-Correa, Sturaro, Valdujo, Haddad, Grant, Faivovich, Lemmon, and Lemmon, 2025, Mol. Phylogenet. Evol., 204 (108275): 1–18.

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist