Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dendropsophus timbeba (Martins and Cardoso, 1987)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dendropsophus > Species: Dendropsophus timbeba

Hyla timbeba Martins and Cardoso, 1987, Rev. Brasil. Biol., 47: 550. Holotype: MZUSP 60550, by original designation. Type locality: "caminho para a Vila Boa Vista, Município de Xapuri" (10° 36′ S, 68° 32′ W; 160 m), Estado do Acre, Brazil.

Hyla allenorum Duellman and Trueb, 1989, Herpetologica, 45: 2. Holotype: KU 207606, by original designation. Type locality: "Reserva Cuzco Amazónico, on the Río Madre de Dios, approximately 15 km E of Puerto Maldonado, 200 m (12° 33′ S, 69° 03′ W), Departamento de Madre de Dios, Peru". Synonymy by Orrico, Duellman, Souza, and Haddad, 2013, J. Herpetol., 47: 615-618.

Dendropsophus allenorum — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 93.

Dendropsophus timbeba — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 93.

Common Names

Cardoso's Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

Cuzco Reserve Treefrog (Dendropsophus allenorum [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 54).

Distribution

Acre (state), upper Amazon Basin, southwestern Brazil to southeastern Peru, possibly into adjacent Bolivia.

Geographic Occurrence

Natural Resident: Brazil, Peru

Likely/Controversially Present: Bolivia

Comment

In the Hyla parviceps group according to the original publication and Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 859. Rodríguez and Duellman, 1994, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 22: 23, provided a brief account as Hyla allenorum. De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 57, and Köhler, 2000, Bonn. Zool. Monogr., 48: 69, consider this species possibly to occur in Bolivia. Duellman, 2005, Cusco Amazonico: 196–198, provided (as Hyla allenorum) an account (adult and larval morphology, description of the call, life history). In the Dendropsophus parviceps group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 93. Fouquet, Noonan, Blanc, and Orrico, 2011, Zootaxa, 3035: 59-67, found this species (as Dendropsophus allenorum) to be distantly related to the Dendropsophus parviceps group sensu stricto, and relatively basal within Dendropsophus. See brief account for the Manu region, Peru, by Villacampa-Ortega, Serrano-Rojas, and Whitworth, 2017, Amph. Manu Learning Cent.: 150–151. In the Dendropsophus parviceps group of Orrico, Grant, Faivovich, Rivera-Correa, Rada, Lyra, Cassini, Valdujo, Schargel, Machado, Wheeler, Barrio-Amorós, Loebmann, Moravec, Zina, Solé, Sturaro, Peloso, Suárez, and Haddad, 2021, Cladistics, 37: 73–105.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.