Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Dendropsophus triangulum (Günther, 1869)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Dendropsophus > Species: Dendropsophus triangulum

Hyla triangulum Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 489. Holotype: BMNH 1947.2.23.88 (formerly 1868.11.15.2) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 95. Type locality: "Brazil"; considered to be of unknown provenance by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 388.

Hyla leucophyllata triangulum — Cope, 1870 "1869", Proc. Am. Philos. Soc., 11: 155.

Dendropsophus triangulum — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 91.

Hyla favosa Cope, 1885, Proc. Am. Philos. Soc., 23: 95. Holotype: ANSP 11483, according to Malnate, 1971, Proc. Acad. Nat. Sci. Philadelphia, 123: 350. Type locality: "Pebas, Upper Amazon" Department of Loreto, Peru. Synonymy with Hyla leucophyllata by Titus, Hillis, and Duellman, 1989, Herpetologica, 45: 17-23; with Dendropsophus triangulum by Caminer, Milá, Jansen, Fouquet, Venegas, Chávez, Lougheed, and Ron, 2017, PLoS One, 12(3: e0171785): 18.

English Names

Triangle Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

Distribution

Amazon basin of Ecuador (Orellana and Sucumbíos provinces), Peru (regions of Loreto and Cusco) and Brazil (states of Acre, Amazonas and Pará), 34 to 387 m elevation. See comment regarding cryptic associates in Colombia and Brazil.

Comment

See Duellman, 1974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 27: 19. Duellman, 1978, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 65: 142–143, 167–168, provided brief accounts (as Hyla favosa and Hyla triangulum) including characterization of call and tadpole. Many color morphs of this species have been recognized taxonomically (Duellman, 1985, in Frost (ed.), Amph. Species World: 155). Considered by Cope, 1870 "1869", Proc. Am. Philos. Soc., 11: 156, and Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 387, to be a junior synonym of Hyla leucophyllata. See comments by Heyer, 1977, Pap. Avulsos Zool., São Paulo, 31: 144. Rodríguez and Duellman, 1994, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 22: 38, provided a brief account for the Iquitos region of northeastern Peru as Hyla triangulum. In the Dendropsophus leucophyllatus group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 91 Duellman, 2005, Cusco Amazonico: 215–218, provided (as Hyla leucophyllata) an account (adult and larval morphology, description of the call, life history). França and Venâncio, 2010, Biotemas, 23: 71–84, provided a record for the municipality of Boca do Acre, Amazonas, with a brief discussion of the range. Bernarde, Machado, and Turci, 2011, Biota Neotrop., 11: 117–144, reported specimens from Reserva Extrativista Riozinho da Liberdade, Acre, Brazil. Lynch and Suárez-Mayorga, 2011, Caldasia, 33: 235–270, illustrated the tadpole and included the species in a key to the tadpoles of Amazonian Colombia. Caminer, Milá, Jansen, Fouquet, Venegas, Chávez, Lougheed, and Ron, 2017, PLoS One, 12(3: e0171785): 1–42, revised the group, provided a detailed account, phylogenetic placement, and redelimited the species, noting that, based on acoustic data and molecular markers, pre-revision Dendropsophus triangulum is a complex of nine candidate species, several of which remain unnamed. Zimmerman, 1983, Herpetologica, 39: 235–246, reported on advertisement call, as Hyla triangulum. See brief account for the Manu region, Peru, by Villacampa-Ortega, Serrano-Rojas, and Whitworth, 2017, Amph. Manu Learning Cent.: 152–153. Metcalf, Marsh, Torres Pacaya, Graham, and Gunnels, 2020, Herpetol. Notes, 13: 753–767, reported the species from the Santa Cruz Forest Reserve, Loreto, northeastern Peru. In the Dendropsophus leucophyllatus group of Orrico, Grant, Faivovich, Rivera-Correa, Rada, Lyra, Cassini, Valdujo, Schargel, Machado, Wheeler, Barrio-Amorós, Loebmann, Moravec, Zina, Solé, Sturaro, Peloso, Suárez, and Haddad, 2021, Cladistics, 37: 73–105.

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For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.