Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Ecnomiohyla miliaria (Cope, 1886)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Ecnomiohyla > Species: Ecnomiohyla miliaria

Hypsiboas miliarius Cope, 1886, Proc. Am. Philos. Soc., 23: 272. Holotype: USNM 14193 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 62. Type locality: "Nicaragua".

Hyla miliaria — Günther, 1901, Biol. Centr. Amer., Rept. Batr., Vol. 7, Part 166: 283; Taylor, 1949, Am. Mus. Novit., 1437: 1–21.

Plectrohyla miliaria — Taylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 596.

Hyla immensa Taylor, 1952, Univ. Kansas Sci. Bull., 35: 815. Holotype: KU 30404, by original designation. Type locality: "Inter-American Institute of Agricultural Sciences, [2 km east of] Turrialba, [Provincia Cartago,] Costa Rica". Savage, 1974, Rev. Biol. Tropical, 22: 94, commented on the type locality. Synonymy by Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 352.

Ecnomiohyla miliaria — Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 100.

English Names

Cope's Brown Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 56).

Distribution

Humid lowlands and premontane slopes from eastern Honduras and southeastern Nicaragua to southeastern Costa Rica on the Atlantic versant (20–900 m) and on the Pacific versant in humid premontane areas of southwestern Costa Rica and western Panama, 1000–1300 m elevation; isolated record from El Salto, municipality of Buenaventura, Valle del Cauca department, Colombia, 750 m elevation.

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Honduras, Nicaragua, Panama

Comment

See Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 352–355; Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 899-901; Savage, 2002, Amph. Rept. Costa Rica: 334–335. Köhler, 1999, Cour. Forschungsinst. Senckenb., 213: 21, reported the species for Nicaragua. McCranie, Townsend, and Wilson, 2003, Caribb. J. Sci., 39: 398–399, reported the species in Honduras. McCranie, 2007, Herpetol. Rev., 38: 38, summarized the departmental distribution in Honduras. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 245. See comments by Sunyer, Páiz, Dehling, and Köhler, 2009, Herpetol. Notes, 2: 189–202, regarding Nicaraguan populations. Köhler, 2011, Amph. Cent. Am.: 222–226, provided a brief summary of natural history for the species of Ecnomiohyla in Central America and provided a range map and photograph for this species. Valencia-Zuleta, Jaramillo-Martinez, Cardona-Botero, and Castro-Herrera, 2016, Herpetol. Notes, 9: 23–26, provided a record for the Colombian lowlands (Valle del Cauca Department). Díaz Gómez, Sunyer, Loza-Molina, Fernández, Salazar-Saavedra, and Martínez-Fonseca, 2017, Mesoam. Herpetol., 4: 658–659, provided a record from southeastern Nicaragua on the Río San Juan. Barrio-Amorós and Torres, 2017, Mesoam. Herpetol., 4: 660–661, provided a record for the Pacific versant of Costa Rica.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.