Ecnomiohyla miliaria (Cope, 1886)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Ecnomiohyla > Species: Ecnomiohyla miliaria

Hypsiboas miliarius Cope, 1886, Proc. Am. Philos. Soc., 23: 272. Holotype: USNM 14193 according to Cochran, 1961, Bull. U.S. Natl. Mus., 220: 62. Type locality: "Nicaragua".

Hyla miliariaGünther, 1901, Biol. Centr. Amer., Rept. Batr., Vol. 7, Part 166: 283; Taylor, 1949, Am. Mus. Novit., 1437: 1–21.

Plectrohyla miliariaTaylor and Smith, 1945, Proc. U.S. Natl. Mus., 95: 596.

Hyla immensa Taylor, 1952, Univ. Kansas Sci. Bull., 35: 815. Holotype: KU 30404, by original designation. Type locality: "Inter-American Institute of Agricultural Sciences, [2 km east of] Turrialba, [Provincia Cartago,] Costa Rica". Savage, 1974, Rev. Biol. Tropical, 22: 94, commented on the type locality. Synonymy by Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 352.

Ecnomiohyla miliariaFaivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 100.

English Names

Cope's Brown Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 56).


Humid lowlands and premontane slopes from eastern Honduras and southeastern Nicaragua to southeastern Costa Rica on the Atlantic versant (20–900 m) and on the Pacific versant in humid premontane areas of southwestern Costa Rica and western Panama, 1000–1300 m elevation; isolated record from El Salto, municipality of Buenaventura, Valle del Cauca department, Colombia, 750 m elevation. 

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Honduras, Nicaragua, Panama


See Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 352–355; Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 899-901; Savage, 2002, Amph. Rept. Costa Rica: 334–335. Köhler, 1999, Cour. Forschungsinst. Senckenb., 213: 21, reported the species for Nicaragua. McCranie, Townsend, and Wilson, 2003, Caribb. J. Sci., 39: 398–399, reported the species in Honduras. McCranie, 2007, Herpetol. Rev., 38: 38, summarized the departmental distribution in Honduras. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 245. See comments by Sunyer, Páiz, Dehling, and Köhler, 2009, Herpetol. Notes, 2: 189–202, regarding Nicaraguan populations. Köhler, 2011, Amph. Cent. Am.: 222–226, provided a brief summary of natural history for the species of Ecnomiohyla in Central America and provided a range map and photograph for this species. Valencia-Zuleta, Jaramillo-Martinez, Cardona-Botero, and Castro-Herrera, 2016, Herpetol. Notes, 9: 23–26, provided a record for the Colombian lowlands (Valle del Cauca Department). Díaz Gómez, Sunyer, Loza-Molina, Fernández, Salazar-Saavedra, and Martínez-Fonseca, 2017, Mesoam. Herpetol., 4: 658–659, provided a record from southeastern Nicaragua on the Río San Juan. Barrio-Amorós and Torres, 2017, Mesoam. Herpetol., 4: 660–661, provided a record for the Pacific versant of Costa Rica. 

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