Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyla sarda (De Betta, 1853)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Hyla > Species: Hyla sarda

Dendrohyas sarda De Betta, 1853, Mem. Accad. Agric. Commerc. Arti Verona, 35: 284. Type(s): Not stated or known to exist. Type locality: "Sardegna", Italy. Manuscript name of Bonelli stated to be a variety of Hyla viridis.

Hyla arborea savignyi var. fuscomaculata Camerano, 1884 "1883", Mem. Accad. Sci. Torino, Ser. 2, 35: 127. Types: MZUT. Type locality: "Sardegna", Italy. Unavailable tetranominal.

Hyla arborea sarda — Pavlov, 1933, Publ. Mus. Hoang Ho Pai Ho, Tien-Tsin, 23: 6. clearly based on misidentified specimens from Manchuria—DRF; Mertens, 1955, Senckenb. Biol., 36: 288; Schneider, 1974, Oecologia, Berlin, 14: 110.

Hyla sarda — Lanza, 1983, Biogeographia, 8: 730.

Hyla (Hyla) sarda — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331, by implication.

Common Names

Sardinia Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

Sardinian Treefrog (Stumpel-Rienks, 1992, Ergänzungsband Handbuch Rept. Amph. Eur., Trivialnamen der Herpetofauna Eur.: 50).

Tyrrhenian Tree Frog (Arnold, 2002, Rept. Amph. Eur., Ed. 2: 78).

Distribution

Sardinia (Italy), Corsica (France), and nearby islands in the Tyrrhenian Sea.

Geographic Occurrence

Natural Resident: France, Italy

Comment

Elevated from status as a subspecies of Hyla arborea by Lanza, 1983, Biogeographia, 8: 730. See account of Corsican population by Delaguerre and Cheylan, 1992, Atlas Repart. Batr. Rept. Corse: 35-37. Lanza, 1983, Guide Reconoscimento Spec. Animali, Anf. Rett.: 115-116, reported on the Italian populations. Disi, Modry, Necas, and Rifai, 2001, Amph. Rept. Hashemite Kingdom Jordan: 97-98, provided an account for the Jordanian population. Arnold, 2002, Rept. Amph. Eur., Ed. 2: 78-79, provided a brief account, figure, and map. Nöllert and Nöllert, 1992, Die Amph. Eur.: 310-312, provided an account and polygon map. In the Hyla arborea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102. Rosso, Castellano, and Giacoma, 2004, Ital. J. Zool., 71: 169–173, reported on the advertisement call. Stöck, Dubey, Klütsch, Litvinchuk, Scheidt, and Perrin, 2008, Mol. Phylogenet. Evol., 49: 1019-1024, reported on the molecular phylogenetic relationships of the species. See detailed account by Corti, 2007, in Lanza et al. (eds.), Fauna d'Italia, 42 (Amph.): 346-351. Bisconti, Canestrelli, and Nascetti, 2011, Biol. J. Linn. Soc., 103: 159-167, reported on genetic variation as evidenced by allozyme electrophoresis. Grosse, 2011, Laubfrösche: 1-286, reviewed the biology of the species. Cadeddu, Giacoma, and Castellano, 2012, J. Zool., London, 286: 285-292, discussed sexual dimorphism. Gvoždík, Canestrelli, García-París, Moravec, Nascetti, Recuero, Teixeira, and Kotlík, 2015, Mol. Phylogenet. Evol., 83: 143–155, reported on molecular phylogenetics and biogeography. Stöck, Dufresnes, Litvinchuk, Lymberakis, Biollay, Berroneau, Borzée, Ghali, Ogielska, and Perrin, 2012, Mol. Phylogenet. Evol., 65: 1–9, reported on the genetic diversity among populations of the Hyla arborea group and provided a dot map of the range, including this species. Speybroeck, Beukema, Bok, and Van Der Voort, 2016, Field Guide Amph. Rept. Brit. Eur.: 161–162, provided a brief account and range map. See Dufresnes, 2019, Amph. Eur., N. Afr., & Middle East: 66, for brief summary of identifying morphology and biology, a range map, as well as a photograph. Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi, and Bassu, 2022, Acta Herpetol., Firenze, 17: 125–133, reported on the range in Sardinia.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.