Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyloscirtus tolkieni Sánchez-Nivicela, Falcón-Reibán, and Cisneros-Heredia, 2023

Class: Amphibia > Order: Anura > Family: Hylidae > Genus: Hyloscirtus > Species: Hyloscirtus tolkieni

Hyloscirtus tolkieni Sánchez-Nivicela, Falcón-Reibán, and Cisneros-Heredia, 2023, ZooKeys, 1141: 79. Holotype: ZSFQ-4142, by original designation. Type locality: "Puente de Piedra (2°47'13"S, 78°36'16"W; 3190 m), Parque Nacional Río Negro- Sopladora, provincia de Morona Santiago, República del Ecuador". https://zoobank.org/0DA4A78A-D514-43FA-B5DA-8F5074F9E353

Common Names

Río Negro Stream Treefrog (original publication).

Tolkien's Stream Treefrog (Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: 452).

Torrentícola de Tolkien (Spanish: Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: 452).

Distribution

Known from the Río Negro-Sopladora National Park, province of Morona Santiago, 3190 m elevation, and 8.4 km south in the same mountain range at Santa Teresita (3070 m elevation), cantón El Pan, Azuay Province, Ecuador,

Geographic Occurrence

Natural Resident: Ecuador

Endemic: Ecuador

Comment

Likely in the Hyloscirtus larinopygion group, according to the original publication, where comparative morphology of the adult was detailed. Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: 452–453, provided an account, with photographs, which summarized identification, adult morphology, systematics, natural history, distribution (including a dot map for Ecuador), and conservation. Sánchez-Nivicela, Moya-Mejía, Calderón, and Carrión-Olmedo, 2026, Rev. Latinoam. Herpetol., 9: 417–428, provided a new record from Santa Teresita, cantón El Pan, Azuay Province, Ecuador, in the same mountain range as the type locality. They also provided the advertisement call, and molecular (12S and 16S mtDNA) evidence placing the species firmly in the Hyloscirtus larinopygion group.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador