Nyctimystes gramineus (Boulenger, 1905)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Pelodryadinae > Genus: Nyctimystes > Species: Nyctimystes gramineus

Hyla graminea Boulenger, 1905, Ann. Mag. Nat. Hist., Ser. 7, 16: 183. Holotype: BMNH 1947.2.23.31 (formerly 1905.1.30.29) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 90. Type locality: "Northern British New Guinea, altitude 900 feet".

Litoria gramineaTyler, 1971, Univ. Kansas Publ. Mus. Nat. Hist., 19: 353.

Dryopsophus gramineus — Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 40. 

Ranoidea graminea — Dubois and Frétey, 2016, Dumerilia, 6: 21. 

Nyctimystes gramineus  — Here (16 August 2018). Based on molecular evidence of Duellman, Marion, and Hedges, 2016, Zootaxa, 4104: 39, and Rosauer, Laffan, Crisp, Donnellan, and Cook, 2009, Mol. Ecol., 18: Suppl. data, placing former junior synonym Litoria dux into Nyctimystes.  

English Names

Northern New Guinea Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 59).


Northern versant of the Owen Stanley Mountains, Northern Province, and the Huon Peninsula of Morobe Province, Papua New Guinea. 

Geographic Occurrence

Natural Resident: Papua New Guinea

Endemic: Papua New Guinea


See Tyler, 1968, Zool. Verh., Leiden, 96: 100. In the Litoria aruensis group (= Litoria gracilenta group) according to Tyler and Davies, 1978, Aust. J. Zool., Suppl. Ser., 27 (63): 10. Menzies and Tyler, 2004, Aust. J. Zool., 52: 203, partitioned the Litoria gracilenta group and placed this species in the Litoria chloris group. Richards and Oliver, 2006, Zootaxa, 1295: 41–60, defined the Litoria graminea complex, containing this species, Litoria dux, and Litoria sauroni, and provided an account for Litoria graminea, including a redescription of the holotype, and discussion of the range. Menzies, 2006, Frogs New Guinea & Solomon Is.: 127, provided a brief account. Shea and Kraus, 2007, Zootaxa, 1514: 37–60, commented on the type specimens of Litoria dux Richards and Oliver, 2006. Kraus, 2018, Zootaxa, 4457: 264–284, provided an account and suggested that the original study of Litoria dux rested on mixed samples of Litoria graminea (including the holotype) and Litoria sauroni

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