Cruziohyla calcarifer (Boulenger, 1902)

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Phyllomedusinae > Genus: Cruziohyla > Species: Cruziohyla calcarifer

Agalychnis calcarifer Boulenger, 1902, Ann. Mag. Nat. Hist., Ser. 7, 9: 52. Holotype: BMNH 1947.2.24.22 (formerly 1901.5.27.26) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 86. Type locality: "the Rio Durango, 350 feet", Provincia Esmeraldas, Ecuador.

Phyllomedusa (Agalychnis) calcariferLutz, 1950, Mem. Oswaldo Cruz, Rio de Janeiro, 48: 601.

Phyllomedusa calcariferFunkhouser, 1957, Occas. Pap. Nat. Hist. Mus. Stanford Univ., 5: 24.

Cruziohyla calcariferFaivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 113.

English Names

Splendid Leaf Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 52).


From Esmeraldas Province in northwestern Ecuador, through western Colombia at several reported localities (including Mecana in Chocó and Valle de Cauca) and through Panama to the most southerly part of Costa Rica, where it is found in the Fila Carbon area of Comadre, Limón Province.

Geographic Occurrence

Natural Resident: Colombia, Costa Rica, Ecuador, Panama


With the recognition of Cruziohyla sylviae, the literature of Cruziohyla calcarifer becomes suspect as to whether addressing Cruziohyla sylviae, Cruziohyla calcarifer, or both; see account and revision by Gray, 2018, Zootaxa, 4450: 401–426. See earlier literature of nominal Cruziohyla calcarifer by Myers and Duellman, 1982, Am. Mus. Novit., 2752: 25–26, account by Duellman, 1970, Monogr. Mus. Nat. Hist. Univ. Kansas: 120–124, and note by Duellman, 2001, Hylid Frogs Middle Am., Ed. 2: 843–845. Lips and Savage, 1996, Proc. Biol. Soc. Washington, 109: 17–26, included this species (as Agalychnis calcarifer) in a key to the tadpoles found in Costa Rica. See accounts by Savage, 2002, Amph. Rept. Costa Rica: 279–281, and Faivovich, Haddad, Baêta, Jungfer, Álvares, Brandão, Sheil, Barrientos, Barrio-Amorós, Cruz, and Wheeler, 2010, Cladistics, 26: 235, who implied that this nominal taxon may be a composite of cryptic species (from which Cruziohyla sylviae was subsequently named). Köhler, 2011, Amph. Cent. Am.: 202–203, provided a map and photograph of the species. Gray, Taupp, Denés, Elsner-Gearing, and Bewick, 2021, Herpetol. J., 31: 170–176, described larval ontogeny and morphology. 

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